Bank Of New York Mellon 13FHR Quarterly holdings report by institutional manager | BK 13 May 21

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
10X GENOMICS INC-CLASS A	CL A	88025U109	31,572	174,430	SH		DFND	15	133,395	0	41,035
10X GENOMICS INC-CLASS A	CL A	88025U109	74,904	413,836	SH		DFND	15,22	413,836	0	0
10X GENOMICS INC-CLASS A	CL A	88025U109	11,140	61,547	SH		DFND	22	61,547	0	0
10X GENOMICS INC-CLASS A	CL A	88025U109	15	81	SH		DFND	4	81	0	0
10X GENOMICS INC-CLASS A	CL A	88025U109	1,880	10,385	SH		DFND	42	10,385	0	0
10X GENOMICS INC-CLASS A	CL A	88025U109	23,700	130,939	SH		DFND	48	113,113	0	17,826
10X GENOMICS INC-CLASS A	CL A	88025U109	221	1,220	SH		OTR	49	1,037	183	0
10X GENOMICS INC-CLASS A	CL A	88025U109	1,119	6,181	SH		DFND	49	6,181	0	0
1-800-FLOWERS.COM INC-CL A	CL A	68243Q106	1,951	70,677	SH		DFND	15	62,773	0	7,904
1-800-FLOWERS.COM INC-CL A	CL A	68243Q106	2,047	74,127	SH		DFND	48	74,127	0	0
1-800-FLOWERS.COM INC-CL A	CL A	68243Q106	9	340	SH		DFND	59	340	0	0
1LIFE HEALTHCARE INC	COM	68269G107	94,189	2,410,150	SH		DFND	15,22	2,410,150	0	0
1LIFE HEALTHCARE INC	COM	68269G107	20,905	534,929	SH		DFND	15	437,826	0	97,103
1LIFE HEALTHCARE INC	COM	68269G107	7,313	187,134	SH		DFND	22	187,134	0	0
1LIFE HEALTHCARE INC	COM	68269G107	2,319	59,346	SH		DFND	42	59,346	0	0
1LIFE HEALTHCARE INC	COM	68269G107	11,994	306,913	SH		DFND	48	306,913	0	0
1LIFE HEALTHCARE INC	COM	68269G107	11	290	SH		DFND	49	290	0	0
1ST CONSTITUTION BANCORP	COM	31986N102	211	11,999	SH		DFND	15	6,374	0	5,625
1ST CONSTITUTION BANCORP	COM	31986N102	372	21,119	SH		DFND	48	21,119	0	0
1ST SOURCE CORP	COM	336901103	4,174	87,732	SH		DFND	15	73,087	0	14,645
1ST SOURCE CORP	COM	336901103	1,919	40,332	SH		DFND	48	40,332	0	0
1ST SOURCE CORP	COM	336901103	67	1,400	SH		DFND	49	1,400	0	0
21VIANET GROUP INC-ADR	SPON	90138A103	429	13,291	SH		DFND	15	11,824	0	1,467
21VIANET GROUP INC-ADR	SPON	90138A103	2,417	74,819	SH		DFND	48	74,819	0	0
22ND CENTURY GROUP INC	COM	90137F103	286	86,954	SH		DFND	48	86,954	0	0
2U INC	COM	90214J101	21,701	567,632	SH		DFND	15	535,476	0	32,156
2U INC	COM	90214J101	247	6,455	SH		DFND	22	6,455	0	0
2U INC	COM	90214J101	4,703	123,007	SH		DFND	48	103,207	0	19,800
2U INC	COM	90214J101	179	4,692	SH		DFND	49	4,692	0	0
2U INC	COM	90214J101	430	11,255	SH		OTR	49	11,255	0	0
2U INC	COM	90214J101	579	15,141	SH		DFND	59	14,757	0	384
3D SYSTEMS CORP	COM	88554D205	15,875	578,538	SH		DFND	15	472,273	0	106,265
3D SYSTEMS CORP	COM	88554D205	8,784	320,106	SH		DFND	22	320,106	0	0
3D SYSTEMS CORP	COM	88554D205	164	5,990	SH		DFND	4	5,990	0	0
3D SYSTEMS CORP	COM	88554D205	8,393	305,882	SH		DFND	48	305,882	0	0
3D SYSTEMS CORP	COM	88554D205	352	12,814	SH		DFND	49	12,514	0	300
3D SYSTEMS CORP	COM	88554D205	0	1	SH		DFND		1	0	0
3M CO	COM	88579Y101	4,070	21,124	SH		OTR	15,119	21,124	0	0
3M CO	COM	88579Y101	10,051	52,166	SH		DFND	15,22	52,166	0	0
3M CO	COM	88579Y101	3,769	19,561	SH		OTR	15,118	19,561	0	0
3M CO	COM	88579Y101	736,429	3,822,032	SH		DFND	15	2,853,869	0	968,163
3M CO	COM	88579Y101	30,773	159,710	SH		DFND	22	159,710	0	0
3M CO	COM	88579Y101	3,968	20,594	SH		DFND	4	20,594	0	0
3M CO	COM	88579Y101	496	2,572	SH		OTR	4	1,516	1,056	0
3M CO	COM	88579Y101	329,847	1,711,892	SH		DFND	48	1,539,029	0	172,863
3M CO	COM	88579Y101	84,667	439,419	SH		DFND	49	427,000	500	11,919
3M CO	COM	88579Y101	49,763	258,268	SH		OTR	49	83,661	63,002	111,605
3M CO	COM	88579Y101	3,064	15,903	SH		DFND	59	15,699	0	204
4D MOLECULAR THERAPEUTICS IN	COM	35104E100	77	1,780	SH		DFND	15	335	0	1,445
4D MOLECULAR THERAPEUTICS IN	COM	35104E100	808	18,637	SH		DFND	48	18,637	0	0
51JOB INC-ADR	SPON	316827104	2,792	44,597	SH		DFND	15	43,712	0	885
51JOB INC-ADR	SPON	316827104	181	2,899	SH		DFND	40	2,899	0	0
51JOB INC-ADR	SPON	316827104	1,440	23,004	SH		DFND	48	23,004	0	0
51JOB INC-ADR	SPON	316827104	3	40	SH		OTR	49	40	0	0
51JOB INC-ADR	SPON	316827104	0	1	SH		DFND		1	0	0
89BIO INC	COM	282559103	29	1,239	SH		DFND	15	759	0	480
89BIO INC	COM	282559103	479	20,219	SH		DFND	48	20,219	0	0
8X8 INC	COM	282914100	14,623	450,769	SH		DFND	15	413,390	0	37,379
8X8 INC	COM	282914100	8,952	275,953	SH		DFND	22	275,953	0	0
8X8 INC	COM	282914100	28	850	SH		DFND	4	850	0	0
8X8 INC	COM	282914100	8,814	271,689	SH		DFND	48	271,689	0	0
8X8 INC	COM	282914100	1,514	46,661	SH		DFND	49	46,661	0	0
8X8 INC	COM	282914100	0	1	SH		DFND		1	0	0
9 METERS BIOPHARMA INC	COM	654405109	68	58,176	SH		DFND	48	58,176	0	0
908 DEVICES INC	COM	65443P102	171	3,536	SH		DFND	15	2,219	0	1,317
908 DEVICES INC	COM	65443P102	441	9,085	SH		OTR	22,78	9,085	0	0
908 DEVICES INC	COM	65443P102	808	16,654	SH		DFND	48	16,654	0	0
A H BELO CORP-A	COM	001282102	30	14,280	SH		DFND	48	14,280	0	0
A H BELO CORP-A	COM	001282102	0	1	SH		DFND		1	0	0
A10 NETWORKS INC	COM	002121101	5,287	550,126	SH		DFND	15	379,369	0	170,757
A10 NETWORKS INC	COM	002121101	4,420	459,914	SH		OTR	15,66	459,914	0	0
A10 NETWORKS INC	COM	002121101	2,082	216,668	SH		DFND	22	216,668	0	0
A10 NETWORKS INC	COM	002121101	1,905	198,207	SH		DFND	48	198,207	0	0
AAON INC	COM	000360206	14,201	202,836	SH		DFND	15	188,228	0	14,608
AAON INC	COM	000360206	7,399	105,680	SH		DFND	22	105,680	0	0
AAON INC	COM	000360206	2,447	34,957	SH		OTR	22,71	34,957	0	0
AAON INC	COM	000360206	10	140	SH		DFND	4	140	0	0
AAON INC	COM	000360206	7,131	101,855	SH		DFND	48	101,855	0	0
AAON INC	COM	000360206	472	6,749	SH		DFND	49	6,749	0	0
AAON INC	COM	000360206	112	1,605	SH		OTR	49	0	0	1,605
AAR CORP	COM	000361105	6,734	161,671	SH		DFND	15	132,761	0	28,910
AAR CORP	COM	000361105	3,599	86,404	SH		DFND	22	86,404	0	0
AAR CORP	COM	000361105	6	140	SH		DFND	4	140	0	0
AAR CORP	COM	000361105	3,563	85,554	SH		DFND	48	85,554	0	0
AAR CORP	COM	000361105	1,217	29,225	SH		DFND	49	29,225	0	0
AAR CORP	COM	000361105	226	5,421	SH		OTR	49	5,421	0	0
AAR CORP	COM	000361105	3	61	SH		DFND	59	0	0	61
AARON'S CO INC/THE	COM	00258W108	4,241	165,146	SH		DFND	15	136,768	0	28,378
AARON'S CO INC/THE	COM	00258W108	184	7,162	SH		OTR	22,87	7,162	0	0
AARON'S CO INC/THE	COM	00258W108	2,208	85,990	SH		DFND	22	85,990	0	0
AARON'S CO INC/THE	COM	00258W108	4	164	SH		DFND	4	164	0	0
AARON'S CO INC/THE	COM	00258W108	2,219	86,415	SH		DFND	48	86,415	0	0
AARON'S CO INC/THE	COM	00258W108	1	32	SH		OTR	49	0	32	0
AARON'S CO INC/THE	COM	00258W108	240	9,336	SH		DFND	49	9,336	0	0
ABB LTD-SPON ADR	SPON	000375204	22	725	SH		DFND	4	725	0	0
ABB LTD-SPON ADR	SPON	000375204	136	4,451	SH		OTR	49	305	146	4,000
ABB LTD-SPON ADR	SPON	000375204	1,124	36,902	SH		DFND	49	36,902	0	0
ABB LTD-SPON ADR	SPON	000375204	6,759	221,825	SH		DFND	59	215,534	0	6,291
ABB LTD-SPON ADR	SPON	000375204	0	1	SH		DFND		1	0	0
ABBOTT LABORATORIES	COM	002824100	10,607	88,510	SH		OTR	15,118	88,510	0	0
ABBOTT LABORATORIES	COM	002824100	872,494	7,280,493	SH		DFND	15	4,287,284	0	2,993,209
ABBOTT LABORATORIES	COM	002824100	3,534	29,493	SH		OTR	15,119	29,493	0	0
ABBOTT LABORATORIES	COM	002824100	19,069	159,118	SH		DFND	15,22	159,118	0	0
ABBOTT LABORATORIES	COM	002824100	55,966	467,002	SH		DFND	22	467,002	0	0
ABBOTT LABORATORIES	COM	002824100	1,008	8,411	SH		OTR	4	2,561	5,850	0
ABBOTT LABORATORIES	COM	002824100	8,080	67,427	SH		DFND	4	67,161	0	266
ABBOTT LABORATORIES	COM	002824100	13,205	110,187	SH		OTR	40,48	110,187	0	0
ABBOTT LABORATORIES	COM	002824100	569,570	4,752,753	SH		DFND	40	4,521,071	0	231,682
ABBOTT LABORATORIES	COM	002824100	22,863	190,781	SH		DFND	40,22	190,781	0	0
ABBOTT LABORATORIES	COM	002824100	3,758	31,358	SH		DFND	42	31,358	0	0
ABBOTT LABORATORIES	COM	002824100	627,057	5,232,454	SH		DFND	48	4,704,250	0	528,204
ABBOTT LABORATORIES	COM	002824100	288,089	2,403,947	SH		DFND	49	2,369,014	540	34,393
ABBOTT LABORATORIES	COM	002824100	76,421	637,694	SH		OTR	49	221,242	384,702	31,750
ABBOTT LABORATORIES	COM	002824100	17,874	149,149	SH		DFND	59	141,465	0	7,683
ABBVIE INC	COM	00287Y109	17,148	158,459	SH		DFND	15,22	158,459	0	0
ABBVIE INC	COM	00287Y109	991,278	9,159,841	SH		DFND	15	6,150,355	0	3,009,486
ABBVIE INC	COM	00287Y109	19,442	179,649	SH		OTR	15,118	179,649	0	0
ABBVIE INC	COM	00287Y109	5,190	47,959	SH		OTR	15,119	47,959	0	0
ABBVIE INC	COM	00287Y109	195,961	1,810,764	SH		DFND	22	1,810,764	0	0
ABBVIE INC	COM	00287Y109	1,088	10,050	SH		OTR	4	2,945	7,075	30
ABBVIE INC	COM	00287Y109	9,772	90,297	SH		DFND	4	90,297	0	0
ABBVIE INC	COM	00287Y109	228	2,105	SH		DFND	42	2,105	0	0
ABBVIE INC	COM	00287Y109	574,617	5,309,710	SH		DFND	48	4,763,744	0	545,966
ABBVIE INC	COM	00287Y109	329,530	3,045,003	SH		DFND	49	3,000,879	912	43,212
ABBVIE INC	COM	00287Y109	75,965	701,949	SH		OTR	49	259,464	408,341	34,144
ABBVIE INC	COM	00287Y109	9,289	85,834	SH		DFND	59	83,528	0	2,306
ABCAM PLC-SPON ADR	ADS	000380204	8	434	SH		OTR	49	0	434	0
ABCAM PLC-SPON ADR	ADS	000380204	1,400	72,851	SH		DFND	59	72,851	0	0
ABEONA THERAPEUTICS INC	COM	00289Y107	93	49,381	SH		DFND	15	9,836	0	39,545
ABEONA THERAPEUTICS INC	COM	00289Y107	423	224,838	SH		DFND	48	224,838	0	0
ABEONA THERAPEUTICS INC	COM	00289Y107	0	6	SH		DFND		6	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	13,403	390,635	SH		DFND	15	335,686	0	54,949
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	206	6,000	SH		OTR	15,119	6,000	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	5,567	162,242	SH		DFND	22	162,242	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	31	910	SH		DFND	4	910	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	6,091	177,531	SH		DFND	48	177,531	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	1,145	33,371	SH		DFND	49	33,371	0	0
ABERCROMBIE & FITCH CO-CL A	CL A	002896207	5	157	SH		OTR	49	6	151	0
ABIOMED INC	COM	003654100	61,992	194,496	SH		DFND	15	115,403	0	79,093
ABIOMED INC	COM	003654100	499	1,567	SH		OTR	15,118	1,567	0	0
ABIOMED INC	COM	003654100	41	129	SH		OTR	15,119	129	0	0
ABIOMED INC	COM	003654100	78,923	247,617	SH		DFND	15,22	247,617	0	0
ABIOMED INC	COM	003654100	8,782	27,554	SH		OTR	22,71	27,554	0	0
ABIOMED INC	COM	003654100	12,764	40,047	SH		DFND	22	40,047	0	0
ABIOMED INC	COM	003654100	132	413	SH		DFND	4	413	0	0
ABIOMED INC	COM	003654100	1,892	5,936	SH		DFND	42	5,936	0	0
ABIOMED INC	COM	003654100	46,471	145,802	SH		DFND	48	131,965	0	13,837
ABIOMED INC	COM	003654100	3,588	11,256	SH		DFND	49	11,247	0	9
ABIOMED INC	COM	003654100	1,272	3,990	SH		OTR	49	791	3,199	0
ABIOMED INC	COM	003654100	2,603	8,167	SH		DFND	59	8,117	0	50
ABIOMED INC	COM	003654100	0	1	SH		DFND		1	0	0
ABM INDUSTRIES INC	COM	000957100	186	3,650	SH		OTR	15,119	3,650	0	0
ABM INDUSTRIES INC	COM	000957100	29,168	571,809	SH		DFND	15	511,475	0	60,334
ABM INDUSTRIES INC	COM	000957100	1,387	27,190	SH		OTR	22,83	27,190	0	0
ABM INDUSTRIES INC	COM	000957100	8,750	171,542	SH		DFND	22	171,542	0	0
ABM INDUSTRIES INC	COM	000957100	30	595	SH		DFND	4	595	0	0
ABM INDUSTRIES INC	COM	000957100	8,542	167,459	SH		DFND	48	167,459	0	0
ABM INDUSTRIES INC	COM	000957100	1,816	35,592	SH		DFND	49	35,592	0	0
ACACIA RESEARCH CORP	ACAC	003881307	436	65,517	SH		DFND	15	18,595	0	46,922
ACACIA RESEARCH CORP	ACAC	003881307	746	112,242	SH		DFND	48	112,242	0	0
ACACIA RESEARCH CORP	ACAC	003881307	34	5,158	SH		DFND	49	5,158	0	0
ACACIA RESEARCH CORP	ACAC	003881307	0	1	SH		DFND		1	0	0
ACADEMY SPORTS & OUTDOORS IN	COM	00402L107	589	21,830	SH		DFND	15	6,381	0	15,449
ACADEMY SPORTS & OUTDOORS IN	COM	00402L107	1,690	62,623	SH		DFND	48	62,623	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	273	4,772	SH		OTR	15,119	4,772	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	297	5,200	SH		DFND	15,22	5,200	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	50,092	876,654	SH		DFND	15	710,617	0	166,037
ACADIA HEALTHCARE CO INC	COM	00404A109	10,076	176,337	SH		OTR	15,66	176,337	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	33,174	580,568	SH		DFND	22	580,568	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	1,307	22,871	SH		OTR	22,69	22,871	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	84	1,465	SH		DFND	4	1,465	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	23,161	405,340	SH		DFND	48	378,868	0	26,472
ACADIA HEALTHCARE CO INC	COM	00404A109	2,603	45,552	SH		DFND	49	45,382	0	170
ACADIA HEALTHCARE CO INC	COM	00404A109	481	8,413	SH		OTR	49	8,413	0	0
ACADIA HEALTHCARE CO INC	COM	00404A109	1	22	SH		DFND	59	22	0	0
ACADIA PHARMACEUTICALS INC	COM	004225108	4,551	176,388	SH		DFND	15	123,439	0	52,949
ACADIA PHARMACEUTICALS INC	COM	004225108	609	23,597	SH		OTR	22,87	23,597	0	0
ACADIA PHARMACEUTICALS INC	COM	004225108	267	10,355	SH		DFND	22	10,355	0	0
ACADIA PHARMACEUTICALS INC	COM	004225108	5,277	204,541	SH		DFND	48	170,424	0	34,117
ACADIA PHARMACEUTICALS INC	COM	004225108	63	2,437	SH		DFND	49	2,437	0	0
ACADIA PHARMACEUTICALS INC	COM	004225108	341	13,204	SH		DFND	59	13,204	0	0
ACADIA PHARMACEUTICALS INC	COM	004225108	0	1	SH		DFND		1	0	0
ACADIA REALTY TRUST	COM	004239109	9,001	474,471	SH		DFND	15	399,501	0	74,970
ACADIA REALTY TRUST	COM	004239109	4,223	222,593	SH		DFND	22	222,593	0	0
ACADIA REALTY TRUST	COM	004239109	30	1,570	SH		DFND	4	1,570	0	0
ACADIA REALTY TRUST	COM	004239109	5,619	296,211	SH		DFND	48	296,211	0	0
ACADIA REALTY TRUST	COM	004239109	2,093	110,334	SH		DFND	49	110,334	0	0
ACCEL ENTERTAINMENT INC	COM	00436Q106	483	44,219	SH		DFND	15	31,685	0	12,534
ACCEL ENTERTAINMENT INC	COM	00436Q106	1,299	118,818	SH		DFND	48	118,818	0	0
ACCEL ENTERTAINMENT INC	COM	00436Q106	0	3	SH		DFND		3	0	0
ACCELERATE DIAGNOSTICS INC	COM	00430H102	321	38,589	SH		DFND	15	28,170	0	10,419
ACCELERATE DIAGNOSTICS INC	COM	00430H102	676	81,250	SH		DFND	48	81,250	0	0
ACCELERON PHARMA INC	COM	00434H108	33,817	249,371	SH		DFND	15,22	249,371	0	0
ACCELERON PHARMA INC	COM	00434H108	19,859	146,439	SH		DFND	15	115,423	0	31,016
ACCELERON PHARMA INC	COM	00434H108	4,627	34,123	SH		DFND	22	34,123	0	0
ACCELERON PHARMA INC	COM	00434H108	3	22	SH		DFND	4	22	0	0
ACCELERON PHARMA INC	COM	00434H108	841	6,201	SH		DFND	42	6,201	0	0
ACCELERON PHARMA INC	COM	00434H108	13,894	102,455	SH		DFND	48	86,496	0	15,959
ACCELERON PHARMA INC	COM	00434H108	20	148	SH		OTR	49	0	148	0
ACCELERON PHARMA INC	COM	00434H108	625	4,611	SH		DFND	49	4,611	0	0
ACCELERON PHARMA INC	COM	00434H108	37	276	SH		DFND	59	217	0	59
ACCENTURE PLC-CL A	SHS	G1151C101	15,702	56,840	SH		DFND	15,22	56,840	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	5,920	21,430	SH		OTR	15,118	21,430	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	730,910	2,645,828	SH		DFND	15	1,570,514	0	1,075,314
ACCENTURE PLC-CL A	SHS	G1151C101	3,403	12,317	SH		OTR	15,119	12,317	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	29,915	108,289	SH		DFND	22	108,289	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	5,210	18,859	SH		DFND	4	18,748	0	111
ACCENTURE PLC-CL A	SHS	G1151C101	3,060	11,078	SH		OTR	4	402	10,359	317
ACCENTURE PLC-CL A	SHS	G1151C101	574,122	2,078,269	SH		DFND	40	1,910,399	0	167,870
ACCENTURE PLC-CL A	SHS	G1151C101	29,314	106,113	SH		DFND	40,22	106,113	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	16,957	61,383	SH		OTR	40,48	61,383	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	3,263	11,810	SH		DFND	42	11,810	0	0
ACCENTURE PLC-CL A	SHS	G1151C101	528,150	1,911,855	SH		DFND	48	1,714,119	0	197,736
ACCENTURE PLC-CL A	SHS	G1151C101	161,574	584,882	SH		DFND	49	562,802	0	22,080
ACCENTURE PLC-CL A	SHS	G1151C101	69,496	251,571	SH		OTR	49	85,273	74,934	91,364
ACCENTURE PLC-CL A	SHS	G1151C101	27,715	100,326	SH		DFND	59	96,836	0	3,490
ACCENTURE PLC-CL A	SHS	G1151C101	667	2,415	SH		DFND		2,415	0	0
ACCO BRANDS CORP	COM	00081T108	6,715	795,577	SH		DFND	15	710,662	0	84,915
ACCO BRANDS CORP	COM	00081T108	1,975	233,978	SH		DFND	48	233,978	0	0
ACCO BRANDS CORP	COM	00081T108	41	4,806	SH		DFND	49	3,487	0	1,319
ACCO BRANDS CORP	COM	00081T108	0	1	SH		DFND		1	0	0
ACCOLADE INC	COM	00437E102	1,194	26,317	SH		DFND	15	17,427	0	8,890
ACCOLADE INC	COM	00437E102	192	4,225	SH		DFND	22	4,225	0	0
ACCOLADE INC	COM	00437E102	3,037	66,942	SH		DFND	48	66,942	0	0
ACCOLADE INC	COM	00437E102	16	353	SH		OTR	49	0	353	0
ACCURAY INC	COM	004397105	537	108,486	SH		DFND	15	82,077	0	26,409
ACCURAY INC	COM	004397105	1,346	272,000	SH		OTR	22,83	272,000	0	0
ACCURAY INC	COM	004397105	1,091	220,428	SH		DFND	48	220,428	0	0
ACCURAY INC	COM	004397105	0	23	SH		DFND		23	0	0
ACELRX PHARMACEUTICALS INC	COM	00444T100	82	48,456	SH		DFND	15	10,589	0	37,867
ACELRX PHARMACEUTICALS INC	COM	00444T100	300	176,483	SH		DFND	48	176,483	0	0
ACER THERAPEUTICS INC	COM	00444P108	40	12,706	SH		DFND	48	12,706	0	0
ACI WORLDWIDE INC	COM	004498101	36	934	SH		OTR	15,119	934	0	0
ACI WORLDWIDE INC	COM	004498101	17,067	448,554	SH		DFND	15	409,817	0	38,737
ACI WORLDWIDE INC	COM	004498101	301	7,900	SH		DFND	15,22	7,900	0	0
ACI WORLDWIDE INC	COM	004498101	5,261	138,273	SH		DFND	22	138,273	0	0
ACI WORLDWIDE INC	COM	004498101	3,282	86,253	SH		OTR	22,87	86,253	0	0
ACI WORLDWIDE INC	COM	004498101	22	585	SH		DFND	4	585	0	0
ACI WORLDWIDE INC	COM	004498101	19,911	523,284	SH		DFND	48	523,284	0	0
ACI WORLDWIDE INC	COM	004498101	262	6,875	SH		OTR	49	3,950	0	2,925
ACI WORLDWIDE INC	COM	004498101	1,477	38,830	SH		DFND	49	38,610	0	220
ACI WORLDWIDE INC	COM	004498101	840	22,064	SH		DFND	59	22,064	0	0
ACLARIS THERAPEUTICS INC	COM	00461U105	1,534	60,889	SH		OTR	22,69	60,889	0	0
ACLARIS THERAPEUTICS INC	COM	00461U105	1,267	50,267	SH		DFND	48	50,267	0	0
ACM RESEARCH INC-CLASS A	COM	00108J109	804	9,949	SH		DFND	15	9,949	0	0
ACM RESEARCH INC-CLASS A	COM	00108J109	514	6,359	SH		DFND	48	6,359	0	0
ACME UNITED CORP	COM	004816104	582	14,753	SH		DFND	15	14,753	0	0
ACME UNITED CORP	COM	004816104	59	1,489	SH		DFND	48	1,489	0	0
ACME UNITED CORP	COM	004816104	0	1	SH		DFND		1	0	0
ACNB CORP	COM	000868109	624	21,284	SH		DFND	15	15,879	0	5,405
ACNB CORP	COM	000868109	592	20,217	SH		DFND	48	20,217	0	0
ACNB CORP	COM	000868109	0	1	SH		DFND		1	0	0
ACTINIUM PHARMACEUTICALS INC	COM	00507W206	340	44,630	SH		DFND	15	44,630	0	0
ACTINIUM PHARMACEUTICALS INC	COM	00507W206	47	6,163	SH		DFND	48	6,163	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	6,428	69,115	SH		DFND	15,22	69,115	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	1,032	11,092	SH		OTR	15,119	11,092	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	2,433	26,156	SH		OTR	15,118	26,156	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	398,169	4,281,383	SH		DFND	15	2,883,705	0	1,397,678
ACTIVISION BLIZZARD INC	COM	00507V109	34,262	368,412	SH		DFND	22	368,412	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	385	4,143	SH		DFND	4	4,143	0	0
ACTIVISION BLIZZARD INC	COM	00507V109	212,965	2,289,945	SH		DFND	48	2,056,583	0	233,362
ACTIVISION BLIZZARD INC	COM	00507V109	15,585	167,578	SH		DFND	49	166,638	0	940
ACTIVISION BLIZZARD INC	COM	00507V109	1,263	13,578	SH		OTR	49	3,454	10,124	0
ACTIVISION BLIZZARD INC	COM	00507V109	4,737	50,937	SH		DFND	59	49,766	0	1,171
ACTIVISION BLIZZARD INC	COM	00507V109	48	520	SH		DFND		520	0	0
ACUITY BRANDS INC	COM	00508Y102	290	1,759	SH		OTR	15,119	1,759	0	0
ACUITY BRANDS INC	COM	00508Y102	465	2,820	SH		DFND	15,22	2,820	0	0
ACUITY BRANDS INC	COM	00508Y102	28,383	172,020	SH		DFND	15	154,349	0	17,671
ACUITY BRANDS INC	COM	00508Y102	7,069	42,844	SH		DFND	22	42,844	0	0
ACUITY BRANDS INC	COM	00508Y102	126	765	SH		DFND	4	765	0	0
ACUITY BRANDS INC	COM	00508Y102	56	338	SH		DFND	40	338	0	0
ACUITY BRANDS INC	COM	00508Y102	21,790	132,059	SH		DFND	48	121,116	0	10,943
ACUITY BRANDS INC	COM	00508Y102	3	16	SH		OTR	49	0	16	0
ACUITY BRANDS INC	COM	00508Y102	4,260	25,821	SH		DFND	49	25,740	0	81
ACUITY BRANDS INC	COM	00508Y102	0	1	SH		DFND	59	1	0	0
ACUSHNET HOLDINGS CORP	COM	005098108	6,597	159,612	SH		DFND	15	134,182	0	25,430
ACUSHNET HOLDINGS CORP	COM	005098108	1,505	36,405	SH		OTR	22,121	36,405	0	0
ACUSHNET HOLDINGS CORP	COM	005098108	5,034	121,801	SH		DFND	40	121,801	0	0
ACUSHNET HOLDINGS CORP	COM	005098108	3,408	82,470	SH		DFND	48	82,470	0	0
ACUSHNET HOLDINGS CORP	COM	005098108	29	706	SH		DFND	49	706	0	0
ACUSHNET HOLDINGS CORP	COM	005098108	0	1	SH		DFND		1	0	0
ACUTUS MEDICAL INC	COM	005111109	898	67,128	SH		DFND	15	61,045	0	6,083
ACUTUS MEDICAL INC	COM	005111109	1,007	75,311	SH		DFND	22	75,311	0	0
ACUTUS MEDICAL INC	COM	005111109	82	6,117	SH		OTR	22,78	6,117	0	0
ACUTUS MEDICAL INC	COM	005111109	423	31,641	SH		DFND	48	31,641	0	0
ADAMAS PHARMACEUTICALS INC	COM	00548A106	248	51,670	SH		DFND	15	51,670	0	0
ADAMAS PHARMACEUTICALS INC	COM	00548A106	161	33,509	SH		DFND	48	33,509	0	0
ADAMAS PHARMACEUTICALS INC	COM	00548A106	0	76	SH		DFND	49	76	0	0
ADAMS RESOURCES & ENERGY INC	COM	006351308	676	24,140	SH		DFND	15	22,855	0	1,285
ADAMS RESOURCES & ENERGY INC	COM	006351308	147	5,261	SH		DFND	48	5,261	0	0
ADAPTHEALTH CORP	COM	00653Q102	8,303	225,860	SH		OTR	15,66	225,860	0	0
ADAPTHEALTH CORP	COM	00653Q102	26,125	710,696	SH		DFND	15	510,958	0	199,738
ADAPTHEALTH CORP	COM	00653Q102	20,418	555,451	SH		DFND	22	555,451	0	0
ADAPTHEALTH CORP	COM	00653Q102	10,127	275,498	SH		DFND	48	275,498	0	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	7,439	184,777	SH		DFND	15	134,844	0	49,933
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	18,863	468,541	SH		DFND	15,22	468,541	0	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	1,221	30,324	SH		DFND	22	30,324	0	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	456	11,323	SH		DFND	42	11,323	0	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	6,494	161,296	SH		DFND	48	137,897	0	23,399
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	117	2,900	SH		OTR	49	0	2,900	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	130	3,235	SH		DFND	49	3,235	0	0
ADAPTIVE BIOTECHNOLOGIES	COM	00650F109	370	9,179	SH		DFND	59	9,179	0	0
ADDUS HOMECARE CORP	COM	006739106	7,560	72,284	SH		DFND	15	67,228	0	5,056
ADDUS HOMECARE CORP	COM	006739106	2,078	19,870	SH		OTR	22,114	19,870	0	0
ADDUS HOMECARE CORP	COM	006739106	4,092	39,123	SH		DFND	22	39,123	0	0
ADDUS HOMECARE CORP	COM	006739106	2	22	SH		DFND	4	22	0	0
ADDUS HOMECARE CORP	COM	006739106	3,940	37,672	SH		DFND	48	37,672	0	0
ADDUS HOMECARE CORP	COM	006739106	100	960	SH		DFND	49	960	0	0
ADDUS HOMECARE CORP	COM	006739106	194	1,856	SH		DFND	59	1,770	0	86
ADICET BIO INC	COM	007002108	484	37,030	SH		DFND	15	37,030	0	0
ADICET BIO INC	COM	007002108	76	5,773	SH		DFND	48	5,773	0	0
ADIENT PLC	ORD	G0084W101	18,175	411,209	SH		DFND	15	330,393	0	80,816
ADIENT PLC	ORD	G0084W101	493	11,162	SH		OTR	15,119	11,162	0	0
ADIENT PLC	ORD	G0084W101	342	7,740	SH		DFND	15,22	7,740	0	0
ADIENT PLC	ORD	G0084W101	4,957	112,159	SH		DFND	22	112,159	0	0
ADIENT PLC	ORD	G0084W101	5	115	SH		DFND	4	115	0	0
ADIENT PLC	ORD	G0084W101	18,331	414,738	SH		DFND	48	414,738	0	0
ADIENT PLC	ORD	G0084W101	950	21,493	SH		DFND	49	21,232	0	261
ADIENT PLC	ORD	G0084W101	17	388	SH		OTR	49	87	301	0
ADIENT PLC	ORD	G0084W101	0	1	SH		DFND	59	1	0	0
ADIENT PLC	ORD	G0084W101	0	1	SH		DFND		1	0	0
ADMA BIOLOGICS INC	COM	000899104	33	18,716	SH		DFND	15	9,789	0	8,927
ADMA BIOLOGICS INC	COM	000899104	329	186,696	SH		DFND	48	186,696	0	0
ADOBE INC	COM	00724F101	20,477	43,075	SH		DFND	15,22	43,075	0	0
ADOBE INC	COM	00724F101	1,592	3,350	SH		OTR	15,119	3,350	0	0
ADOBE INC	COM	00724F101	7,700	16,198	SH		OTR	15,118	16,198	0	0
ADOBE INC	COM	00724F101	1,072,360	2,255,842	SH		DFND	15	1,443,062	0	812,780
ADOBE INC	COM	00724F101	99,040	208,342	SH		DFND	22	208,342	0	0
ADOBE INC	COM	00724F101	9,708	20,423	SH		DFND	4	20,423	0	0
ADOBE INC	COM	00724F101	650	1,367	SH		OTR	4	596	312	459
ADOBE INC	COM	00724F101	73	154	SH		DFND	40	154	0	0
ADOBE INC	COM	00724F101	689,175	1,449,765	SH		DFND	48	1,300,562	0	149,203
ADOBE INC	COM	00724F101	34,914	73,445	SH		OTR	49	50,000	22,877	568
ADOBE INC	COM	00724F101	327,682	689,320	SH		DFND	49	675,914	163	13,243
ADOBE INC	COM	00724F101	22,096	46,482	SH		OTR	51,48	46,482	0	0
ADOBE INC	COM	00724F101	174,048	366,131	SH		OTR	51,94	366,131	0	0
ADOBE INC	COM	00724F101	49,629	104,400	SH		DFND	51,22	104,400	0	0
ADOBE INC	COM	00724F101	1,551,354	3,263,467	SH		DFND	51	2,233,216	0	1,030,251
ADOBE INC	COM	00724F101	4,564	9,600	SH		OTR	51	9,600	0	0
ADOBE INC	COM	00724F101	10,990	23,118	SH		DFND	59	22,715	0	403
ADT INC	COM	00090Q103	28,594	3,387,863	SH		DFND	15	3,044,822	0	343,041
ADT INC	COM	00090Q103	339	40,181	SH		DFND	22	40,181	0	0
ADT INC	COM	00090Q103	3,203	379,504	SH		DFND	48	333,045	0	46,459
ADT INC	COM	00090Q103	328	38,826	SH		DFND	49	38,826	0	0
ADT INC	COM	00090Q103	1	92	SH		DFND		92	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	16	399	SH		OTR	15,119	399	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	276	6,970	SH		DFND	15,22	6,970	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	11,804	298,531	SH		DFND	15	253,305	0	45,226
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	2,529	63,962	SH		DFND	22	63,962	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	2	50	SH		DFND	4	50	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	8,765	221,673	SH		DFND	48	221,673	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	442	11,167	SH		DFND	49	11,167	0	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	3	67	SH		OTR	49	0	67	0
ADTALEM GLOBAL EDUCATION INC	COM	00737L103	0	1	SH		DFND		1	0	0
ADTRAN INC	COM	00738A106	7,747	464,446	SH		OTR	15,66	464,446	0	0
ADTRAN INC	COM	00738A106	17,070	1,023,411	SH		DFND	15	832,388	0	191,023
ADTRAN INC	COM	00738A106	22,883	1,371,906	SH		DFND	22	1,371,906	0	0
ADTRAN INC	COM	00738A106	12	730	SH		DFND	4	730	0	0
ADTRAN INC	COM	00738A106	4,009	240,357	SH		DFND	48	240,357	0	0
ADTRAN INC	COM	00738A106	771	46,235	SH		DFND	49	46,235	0	0
ADTRAN INC	COM	00738A106	182	10,920	SH		OTR	49	9,350	1,570	0
ADTRAN INC	COM	00738A106	0	1	SH		DFND		1	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	1,146	6,248	SH		DFND	15,22	6,248	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	51,296	279,560	SH		DFND	15	172,409	0	107,151
ADVANCE AUTO PARTS INC	COM	00751Y106	446	2,433	SH		OTR	15,119	2,433	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	416	2,267	SH		OTR	15,118	2,267	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	3,321	18,097	SH		DFND	22	18,097	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	85	464	SH		DFND	4	464	0	0
ADVANCE AUTO PARTS INC	COM	00751Y106	34,864	190,005	SH		DFND	48	169,664	0	20,341
ADVANCE AUTO PARTS INC	COM	00751Y106	196	1,069	SH		OTR	49	794	275	0
ADVANCE AUTO PARTS INC	COM	00751Y106	1,997	10,882	SH		DFND	49	10,837	0	45
ADVANCE AUTO PARTS INC	COM	00751Y106	388	2,114	SH		DFND	59	2,054	0	60
ADVANCE AUTO PARTS INC	COM	00751Y106	1	6	SH		DFND		6	0	0
ADVANCED DRAINAGE SYSTEMS IN	COM	00790R104	12,468	120,592	SH		DFND	15	101,738	0	18,854
ADVANCED DRAINAGE SYSTEMS IN	COM	00790R104	1,580	15,278	SH		OTR	22,69	15,278	0	0
ADVANCED DRAINAGE SYSTEMS IN	COM	00790R104	10,039	97,095	SH		OTR	22,71	97,095	0	0
ADVANCED DRAINAGE SYSTEMS IN	COM	00790R104	13,981	135,227	SH		DFND	48	135,227	0	0
ADVANCED DRAINAGE SYSTEMS IN	COM	00790R104	216	2,085	SH		DFND	49	2,085	0	0
ADVANCED EMISSIONS SOLUTIONS	COM	00770C101	427	77,653	SH		DFND	15	68,014	0	9,639
ADVANCED EMISSIONS SOLUTIONS	COM	00770C101	218	39,577	SH		DFND	48	39,577	0	0
ADVANCED EMISSIONS SOLUTIONS	COM	00770C101	0	1	SH		DFND		1	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	19,317	176,946	SH		DFND	15	163,838	0	13,108
ADVANCED ENERGY INDUSTRIES	COM	007973100	607	5,556	SH		OTR	22,78	5,556	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	10,777	98,721	SH		DFND	22	98,721	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	3,517	32,216	SH		OTR	22,114	32,216	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	58	535	SH		DFND	4	535	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	10,853	99,418	SH		DFND	48	99,418	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	60	550	SH		OTR	49	0	550	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	2,307	21,128	SH		DFND	49	21,128	0	0
ADVANCED ENERGY INDUSTRIES	COM	007973100	0	1	SH		DFND		1	0	0
ADVANCED MICRO DEVICES	COM	007903107	3,212	40,923	SH		OTR	15,118	40,923	0	0
ADVANCED MICRO DEVICES	COM	007903107	402,151	5,122,942	SH		DFND	15	3,082,729	0	2,040,213
ADVANCED MICRO DEVICES	COM	007903107	515	6,562	SH		OTR	15,119	6,562	0	0
ADVANCED MICRO DEVICES	COM	007903107	8,540	108,789	SH		DFND	15,22	108,789	0	0
ADVANCED MICRO DEVICES	COM	007903107	38,999	496,797	SH		DFND	22	496,797	0	0
ADVANCED MICRO DEVICES	COM	007903107	255	3,243	SH		OTR	4	1,450	1,793	0
ADVANCED MICRO DEVICES	COM	007903107	3,784	48,204	SH		DFND	4	48,204	0	0
ADVANCED MICRO DEVICES	COM	007903107	283,080	3,606,114	SH		DFND	48	3,235,048	0	371,066
ADVANCED MICRO DEVICES	COM	007903107	142,077	1,809,895	SH		DFND	49	1,790,090	1,290	18,515
ADVANCED MICRO DEVICES	COM	007903107	4,881	62,175	SH		OTR	49	36,873	25,232	70
ADVANCED MICRO DEVICES	COM	007903107	4,743	60,425	SH		DFND	59	59,752	0	673
ADVANSIX INC	COM	00773T101	3,986	148,606	SH		DFND	15	121,822	0	26,784
ADVANSIX INC	COM	00773T101	1,901	70,865	SH		DFND	22	70,865	0	0
ADVANSIX INC	COM	00773T101	1,904	70,993	SH		DFND	48	70,993	0	0
ADVANSIX INC	COM	00773T101	203	7,576	SH		DFND	49	7,576	0	0
ADVANSIX INC	COM	00773T101	5	182	SH		OTR	49	160	22	0
ADVANSIX INC	COM	00773T101	2	65	SH		DFND	59	65	0	0
ADVANTAGE SOLUTIONS INC	COM	00791N102	657	55,626	SH		DFND	48	55,626	0	0
ADVAXIS INC	COM	007624307	49	66,387	SH		DFND	48	66,387	0	0
ADVENT TECHNOLOGIES HOLDINGS	COM	00788A105	178	13,275	SH		DFND	48	13,275	0	0
ADVERUM BIOTECHNOLOGIES INC	COM	00773U108	980	99,411	SH		DFND	15	66,438	0	32,973
ADVERUM BIOTECHNOLOGIES INC	COM	00773U108	2,199	223,026	SH		DFND	48	223,026	0	0
ADVERUM BIOTECHNOLOGIES INC	COM	00773U108	1	85	SH		DFND	49	85	0	0
AECOM	COM	00766T100	801	12,497	SH		OTR	15,119	12,497	0	0
AECOM	COM	00766T100	40,784	636,153	SH		DFND	15	569,301	0	66,852
AECOM	COM	00766T100	11,252	175,504	SH		DFND	22	175,504	0	0
AECOM	COM	00766T100	141	2,195	SH		DFND	4	2,195	0	0
AECOM	COM	00766T100	35,027	546,364	SH		DFND	48	503,314	0	43,050
AECOM	COM	00766T100	5,244	81,795	SH		DFND	49	81,493	0	302
AECOM	COM	00766T100	24	367	SH		OTR	49	154	213	0
AECOM	COM	00766T100	2	32	SH		DFND	59	32	0	0
AECOM	COM	00766T100	0	1	SH		DFND		1	0	0
AEGION CORP	COM	00770F104	4,421	153,760	SH		DFND	15	128,199	0	25,561
AEGION CORP	COM	00770F104	2,284	79,442	SH		DFND	22	79,442	0	0
AEGION CORP	COM	00770F104	2,229	77,517	SH		DFND	48	77,517	0	0
AEGION CORP	COM	00770F104	273	9,495	SH		DFND	49	9,495	0	0
AEGION CORP	COM	00770F104	0	1	SH		DFND		1	0	0
AEGLEA BIOTHERAPEUTICS INC	COM	00773J103	201	25,383	SH		DFND	15	9,060	0	16,323
AEGLEA BIOTHERAPEUTICS INC	COM	00773J103	927	117,055	SH		DFND	48	117,055	0	0
AEGLEA BIOTHERAPEUTICS INC	COM	00773J103	7	835	SH		DFND	49	835	0	0
AEGON N.V.-NY REG SHR	NY R	007924103	2	450	SH		DFND	4	450	0	0
AEGON N.V.-NY REG SHR	NY R	007924103	8	1,696	SH		OTR	49	0	1,696	0
AEGON N.V.-NY REG SHR	NY R	007924103	168	35,485	SH		DFND	49	35,485	0	0
AEHR TEST SYSTEMS	COM	00760J108	37	14,811	SH		DFND	48	14,811	0	0
AERCAP HOLDINGS NV	SHS	N00985106	8,972	152,746	SH		DFND	15	132,506	0	20,240
AERCAP HOLDINGS NV	SHS	N00985106	41,739	710,564	SH		DFND	15,22	710,564	0	0
AERCAP HOLDINGS NV	SHS	N00985106	913	15,535	SH		OTR	15,119	15,535	0	0
AERCAP HOLDINGS NV	SHS	N00985106	4,114	70,034	SH		DFND	22	70,034	0	0
AERCAP HOLDINGS NV	SHS	N00985106	1,103	18,779	SH		DFND	42	18,779	0	0
AERCAP HOLDINGS NV	SHS	N00985106	2,296	39,079	SH		DFND	48	39,079	0	0
AERCAP HOLDINGS NV	SHS	N00985106	550	9,369	SH		OTR	49	9,300	69	0
AERCAP HOLDINGS NV	SHS	N00985106	18	298	SH		DFND	49	298	0	0
AERCAP HOLDINGS NV	SHS	N00985106	45	759	SH		DFND	59	736	0	23
AERIE PHARMACEUTICALS INC	COM	00771V108	1,080	60,423	SH		DFND	15	42,664	0	17,759
AERIE PHARMACEUTICALS INC	COM	00771V108	924	51,698	SH		OTR	22,87	51,698	0	0
AERIE PHARMACEUTICALS INC	COM	00771V108	1,790	100,155	SH		DFND	48	100,155	0	0
AERIE PHARMACEUTICALS INC	COM	00771V108	4	210	SH		DFND	49	210	0	0
AERIE PHARMACEUTICALS INC	COM	00771V108	24	1,360	SH		OTR	49	0	1,360	0
AEROJET ROCKETDYNE HOLDINGS	COM	007800105	14,187	302,105	SH		DFND	15	276,965	0	25,140
AEROJET ROCKETDYNE HOLDINGS	COM	007800105	8,702	185,302	SH		DFND	22	185,302	0	0
AEROJET ROCKETDYNE HOLDINGS	COM	007800105	79	1,690	SH		DFND	4	1,690	0	0
AEROJET ROCKETDYNE HOLDINGS	COM	007800105	8,815	187,716	SH		DFND	48	187,716	0	0
AEROJET ROCKETDYNE HOLDINGS	COM	007800105	1,480	31,519	SH		DFND	49	31,519	0	0
AEROVIRONMENT INC	COM	008073108	11,505	99,133	SH		OTR	15,66	99,133	0	0
AEROVIRONMENT INC	COM	008073108	21,722	187,161	SH		DFND	15	148,870	0	38,291
AEROVIRONMENT INC	COM	008073108	2,605	22,441	SH		OTR	22,114	22,441	0	0
AEROVIRONMENT INC	COM	008073108	9,924	85,504	SH		DFND	22	85,504	0	0
AEROVIRONMENT INC	COM	008073108	778	6,703	SH		OTR	22,69	6,703	0	0
AEROVIRONMENT INC	COM	008073108	27	230	SH		DFND	4	230	0	0
AEROVIRONMENT INC	COM	008073108	6,401	55,149	SH		DFND	48	55,149	0	0
AEROVIRONMENT INC	COM	008073108	570	4,914	SH		DFND	49	4,914	0	0
AEROVIRONMENT INC	COM	008073108	70	605	SH		OTR	49	0	605	0
AES CORP	COM	00130H105	116,320	4,338,682	SH		DFND	15	3,127,816	0	1,210,866
AES CORP	COM	00130H105	1,669	62,267	SH		DFND	15,22	62,267	0	0
AES CORP	COM	00130H105	902	33,644	SH		OTR	15,119	33,644	0	0
AES CORP	COM	00130H105	5,322	198,517	SH		OTR	15,118	198,517	0	0
AES CORP	COM	00130H105	22,935	855,478	SH		DFND	22	855,478	0	0
AES CORP	COM	00130H105	219	8,173	SH		DFND	4	8,173	0	0
AES CORP	COM	00130H105	3	110	SH		OTR	4	110	0	0
AES CORP	COM	00130H105	132	4,942	SH		DFND	42	4,942	0	0
AES CORP	COM	00130H105	68,137	2,541,475	SH		DFND	48	2,343,588	0	197,887
AES CORP	COM	00130H105	7,970	297,280	SH		DFND	49	294,805	0	2,475
AES CORP	COM	00130H105	336	12,531	SH		OTR	49	7,085	5,356	90
AES CORP	COM	00130H105	737	27,493	SH		DFND	59	27,433	0	60
AFFILIATED MANAGERS GROUP	COM	008252108	305	2,045	SH		OTR	15,119	2,045	0	0
AFFILIATED MANAGERS GROUP	COM	008252108	31,544	211,664	SH		DFND	15	187,416	0	24,248
AFFILIATED MANAGERS GROUP	COM	008252108	137	920	SH		DFND	15,22	920	0	0
AFFILIATED MANAGERS GROUP	COM	008252108	7,004	46,996	SH		DFND	22	46,996	0	0
AFFILIATED MANAGERS GROUP	COM	008252108	92	619	SH		DFND	4	619	0	0
AFFILIATED MANAGERS GROUP	COM	008252108	24,438	163,982	SH		DFND	48	150,320	0	13,662
AFFILIATED MANAGERS GROUP	COM	008252108	16	106	SH		OTR	49	0	106	0
AFFILIATED MANAGERS GROUP	COM	008252108	3,239	21,735	SH		DFND	49	21,597	0	138
AFFILIATED MANAGERS GROUP	COM	008252108	0	2	SH		DFND	59	2	0	0
AFFIMED NV	COM	N01045108	328	41,418	SH		DFND	15	14,517	0	26,901
AFFIMED NV	COM	N01045108	1,010	127,658	SH		OTR	22,69	127,658	0	0
AFFIMED NV	COM	N01045108	1,633	206,489	SH		DFND	48	206,489	0	0
AFFIMED NV	COM	N01045108	5	620	SH		DFND	49	620	0	0
AFFIRM HOLDINGS INC	COM	00827B106	71	997	SH		DFND	15	717	0	280
AFFIRM HOLDINGS INC	COM	00827B106	576	8,139	SH		DFND	15,22	8,139	0	0
AFFIRM HOLDINGS INC	COM	00827B106	58	818	SH		DFND	22	818	0	0
AFFIRM HOLDINGS INC	COM	00827B106	1,166	16,487	SH		DFND	48	16,487	0	0
AFLAC INC	COM	001055102	2,943	57,500	SH		DFND	15,22	57,500	0	0
AFLAC INC	COM	001055102	1,105	21,584	SH		OTR	15,118	21,584	0	0
AFLAC INC	COM	001055102	2,197	42,931	SH		OTR	15,119	42,931	0	0
AFLAC INC	COM	001055102	132,329	2,585,570	SH		DFND	15	1,462,371	0	1,123,199
AFLAC INC	COM	001055102	2,283	44,614	SH		OTR	22,80	44,614	0	0
AFLAC INC	COM	001055102	5,653	110,463	SH		DFND	22	110,463	0	0
AFLAC INC	COM	001055102	54	1,053	SH		OTR	4	0	1,000	53
AFLAC INC	COM	001055102	373	7,290	SH		DFND	4	7,290	0	0
AFLAC INC	COM	001055102	100,342	1,960,571	SH		DFND	48	1,753,019	0	207,552
AFLAC INC	COM	001055102	2,715	53,053	SH		OTR	49	11,259	40,394	1,400
AFLAC INC	COM	001055102	17,921	350,161	SH		DFND	49	348,167	0	1,994
AFLAC INC	COM	001055102	2,346	45,830	SH		DFND	59	44,564	0	1,266
AFLAC INC	COM	001055102	0	1	SH		DFND		1	0	0
AG MORTGAGE INVESTMENT TRUST	COM	001228105	73	18,039	SH		DFND	48	18,039	0	0
AGCO CORP	COM	001084102	51,914	361,390	SH		DFND	15	330,491	0	30,899
AGCO CORP	COM	001084102	398	2,770	SH		OTR	15,119	2,770	0	0
AGCO CORP	COM	001084102	764	5,320	SH		DFND	15,22	5,320	0	0
AGCO CORP	COM	001084102	1,541	10,729	SH		OTR	22,69	10,729	0	0
AGCO CORP	COM	001084102	13,400	93,283	SH		DFND	22	93,283	0	0
AGCO CORP	COM	001084102	141	983	SH		DFND	4	983	0	0
AGCO CORP	COM	001084102	4,814	33,513	SH		DFND	40	33,513	0	0
AGCO CORP	COM	001084102	35,861	249,644	SH		DFND	48	230,876	0	18,768
AGCO CORP	COM	001084102	3,451	24,022	SH		DFND	49	23,706	0	316
AGCO CORP	COM	001084102	11	77	SH		OTR	49	0	77	0
AGCO CORP	COM	001084102	2	12	SH		DFND	59	12	0	0
AGCO CORP	COM	001084102	0	1	SH		DFND		1	0	0
AGEAGLE AERIAL SYSTEMS INC	COM	00848K101	168	26,780	SH		DFND	48	26,780	0	0
AGENUS INC	COM	00847G705	412	151,322	SH		DFND	15	110,068	0	41,254
AGENUS INC	COM	00847G705	1,145	421,001	SH		DFND	48	421,001	0	0
AGENUS INC	COM	00847G705	0	13	SH		DFND		13	0	0
AGILE THERAPEUTICS INC	COM	00847L100	58	27,906	SH		DFND	15	20,394	0	7,512
AGILE THERAPEUTICS INC	COM	00847L100	350	168,043	SH		DFND	48	168,043	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	171,393	1,348,065	SH		DFND	15	855,925	0	492,140
AGILENT TECHNOLOGIES INC	COM	00846U101	3,522	27,699	SH		DFND	15,22	27,699	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	550	4,323	SH		OTR	15,119	4,323	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	1,306	10,274	SH		OTR	15,118	10,274	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	16,014	125,959	SH		DFND	22	125,959	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	371	2,915	SH		DFND	4	2,915	0	0
AGILENT TECHNOLOGIES INC	COM	00846U101	32	249	SH		OTR	4	30	200	19
AGILENT TECHNOLOGIES INC	COM	00846U101	115,721	910,182	SH		DFND	48	817,559	0	92,623
AGILENT TECHNOLOGIES INC	COM	00846U101	27,783	218,525	SH		DFND	49	215,591	0	2,934
AGILENT TECHNOLOGIES INC	COM	00846U101	3,905	30,713	SH		OTR	49	20,065	10,648	0
AGILENT TECHNOLOGIES INC	COM	00846U101	487	3,832	SH		DFND	59	3,832	0	0
AGILYSYS INC	COM	00847J105	4,072	84,914	SH		DFND	15	79,112	0	5,802
AGILYSYS INC	COM	00847J105	2,511	52,354	SH		DFND	22	52,354	0	0
AGILYSYS INC	COM	00847J105	1,532	31,950	SH		OTR	22,86	31,950	0	0
AGILYSYS INC	COM	00847J105	2,765	57,652	SH		DFND	48	57,652	0	0
AGILYSYS INC	COM	00847J105	246	5,139	SH		DFND	49	5,139	0	0
AGILYSYS INC	COM	00847J105	0	1	SH		DFND		1	0	0
AGIOS PHARMACEUTICALS INC	COM	00847X104	5,189	100,490	SH		DFND	15	69,385	0	31,105
AGIOS PHARMACEUTICALS INC	COM	00847X104	257	4,976	SH		DFND	22	4,976	0	0
AGIOS PHARMACEUTICALS INC	COM	00847X104	5,371	104,017	SH		DFND	48	86,406	0	17,611
AGIOS PHARMACEUTICALS INC	COM	00847X104	253	4,893	SH		DFND	49	4,893	0	0
AGIOS PHARMACEUTICALS INC	COM	00847X104	2	42	SH		OTR	49	0	42	0
AGIOS PHARMACEUTICALS INC	COM	00847X104	555	10,744	SH		DFND	59	10,744	0	0
AGIOS PHARMACEUTICALS INC	COM	00847X104	0	1	SH		DFND		1	0	0
AGNC INVESTMENT CORP	COM	00123Q104	581	34,671	SH		OTR	15,119	34,671	0	0
AGNC INVESTMENT CORP	COM	00123Q104	14,645	873,788	SH		DFND	15	596,982	0	276,806
AGNC INVESTMENT CORP	COM	00123Q104	1,042	62,145	SH		DFND	22	62,145	0	0
AGNC INVESTMENT CORP	COM	00123Q104	6	330	SH		OTR	4	330	0	0
AGNC INVESTMENT CORP	COM	00123Q104	315	18,770	SH		DFND	4	18,770	0	0
AGNC INVESTMENT CORP	COM	00123Q104	16,244	969,206	SH		DFND	48	801,785	0	167,421
AGNC INVESTMENT CORP	COM	00123Q104	15,701	936,830	SH		DFND	49	926,760	0	10,070
AGNC INVESTMENT CORP	COM	00123Q104	1,832	109,335	SH		OTR	49	89,043	20,292	0
AGNC INVESTMENT CORP	COM	00123Q104	636	37,924	SH		DFND	59	35,954	0	1,970
AGNICO EAGLE MINES LTD	COM	008474108	120	2,075	SH		OTR	15,119	2,075	0	0
AGNICO EAGLE MINES LTD	COM	008474108	9,014	155,929	SH		DFND	15	126,472	0	29,457
AGNICO EAGLE MINES LTD	COM	008474108	7,204	124,609	SH		DFND	48	124,609	0	0
AGNICO EAGLE MINES LTD	COM	008474108	126	2,182	SH		DFND	49	2,182	0	0
AGNICO EAGLE MINES LTD	COM	008474108	603	10,425	SH		OTR	49	10,300	125	0
AGNICO EAGLE MINES LTD	COM	008474108	151	2,609	SH		DFND	59	2,587	0	22
AGNICO EAGLE MINES LTD	COM	008474108	0	1	SH		DFND		1	0	0
AGORA INC-ADR	ADS	00851L103	12,464	247,944	SH		DFND	15	247,944	0	0
AGREE REALTY CORP	COM	008492100	30,434	452,143	SH		DFND	15	365,257	0	86,886
AGREE REALTY CORP	COM	008492100	7,651	113,674	SH		OTR	15,66	113,674	0	0
AGREE REALTY CORP	COM	008492100	14,782	219,612	SH		DFND	22	219,612	0	0
AGREE REALTY CORP	COM	008492100	36	530	SH		DFND	4	530	0	0
AGREE REALTY CORP	COM	008492100	14,150	210,220	SH		DFND	48	210,220	0	0
AGREE REALTY CORP	COM	008492100	1,671	24,826	SH		DFND	49	24,826	0	0
AGREE REALTY CORP	COM	008492100	1,130	16,788	SH		DFND	59	16,186	0	602
AGROFRESH SOLUTIONS INC	COM	00856G109	40	20,169	SH		DFND	15	3,409	0	16,760
AGROFRESH SOLUTIONS INC	COM	00856G109	148	73,823	SH		DFND	48	73,823	0	0
AIKIDO PHARMA INC	COM	008875106	50	44,190	SH		DFND	48	44,190	0	0
AIKIDO PHARMA INC	COM	008875106	0	1	SH		DFND		1	0	0
AIM IMMUNOTECH INC	COM	00901B105	57	25,266	SH		DFND	48	25,266	0	0
AIM IMMUNOTECH INC	COM	00901B105	0	2	SH		DFND		2	0	0
AIR LEASE CORP	CL A	00912X302	265	5,402	SH		OTR	15,119	5,402	0	0
AIR LEASE CORP	CL A	00912X302	18,253	372,512	SH		DFND	15	320,465	0	52,047
AIR LEASE CORP	CL A	00912X302	409	8,345	SH		DFND	22	8,345	0	0
AIR LEASE CORP	CL A	00912X302	0	10	SH		DFND	4	10	0	0
AIR LEASE CORP	CL A	00912X302	9,060	184,901	SH		DFND	48	152,952	0	31,949
AIR LEASE CORP	CL A	00912X302	1,173	23,942	SH		OTR	49	23,942	0	0
AIR LEASE CORP	CL A	00912X302	95	1,944	SH		DFND	49	1,944	0	0
AIR LEASE CORP	CL A	00912X302	5	95	SH		DFND	59	95	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	369,391	1,312,971	SH		DFND	15	940,925	0	372,046
AIR PRODUCTS & CHEMICALS INC	COM	009158106	5,555	19,745	SH		DFND	15,22	19,745	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	2,101	7,469	SH		OTR	15,118	7,469	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	1,424	5,061	SH		OTR	15,119	5,061	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	34,858	123,901	SH		DFND	22	123,901	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	7,253	25,781	SH		DFND	4	25,781	0	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	3,850	13,686	SH		OTR	4	2,043	11,643	0
AIR PRODUCTS & CHEMICALS INC	COM	009158106	181,642	645,630	SH		DFND	48	578,759	0	66,871
AIR PRODUCTS & CHEMICALS INC	COM	009158106	205,782	731,437	SH		DFND	49	718,219	164	13,054
AIR PRODUCTS & CHEMICALS INC	COM	009158106	27,784	98,757	SH		OTR	49	48,355	49,529	873
AIR PRODUCTS & CHEMICALS INC	COM	009158106	20,662	73,441	SH		DFND	59	69,823	0	3,618
AIR PRODUCTS & CHEMICALS INC	COM	009158106	0	1	SH		DFND		1	0	0
AIR TRANSPORT SERVICES GROUP	COM	00922R105	2,495	85,263	SH		DFND	15	63,939	0	21,324
AIR TRANSPORT SERVICES GROUP	COM	00922R105	2,325	79,460	SH		OTR	22,114	79,460	0	0
AIR TRANSPORT SERVICES GROUP	COM	00922R105	4,186	143,071	SH		DFND	48	143,071	0	0
AIRBNB INC-CLASS A	COM	009066101	7,000	37,248	SH		DFND	15	35,492	0	1,756
AIRBNB INC-CLASS A	COM	009066101	8,740	46,503	SH		DFND	48	46,503	0	0
AIRBNB INC-CLASS A	COM	009066101	483	2,570	SH		OTR	49	2,570	0	0
AIRBNB INC-CLASS A	COM	009066101	373	1,985	SH		DFND	49	1,985	0	0
AIRBNB INC-CLASS A	COM	009066101	233	1,240	SH		DFND	59	1,172	0	68
AKAMAI TECHNOLOGIES INC	COM	00971T101	375	3,684	SH		OTR	15,119	3,684	0	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	68,913	676,278	SH		DFND	15	414,821	0	261,457
AKAMAI TECHNOLOGIES INC	COM	00971T101	559	5,484	SH		OTR	15,118	5,484	0	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	1,524	14,953	SH		DFND	15,22	14,953	0	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	4,411	43,289	SH		DFND	22	43,289	0	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	123	1,207	SH		DFND	4	1,207	0	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	49,144	482,275	SH		DFND	48	433,045	0	49,230
AKAMAI TECHNOLOGIES INC	COM	00971T101	4,443	43,604	SH		DFND	49	43,569	0	35
AKAMAI TECHNOLOGIES INC	COM	00971T101	3,782	37,116	SH		OTR	49	26,506	10,610	0
AKAMAI TECHNOLOGIES INC	COM	00971T101	1,483	14,555	SH		DFND	59	14,320	0	235
AKAMAI TECHNOLOGIES INC	NOTE	00971TAJ0	848	700,000	SH		DFND	15	700,000	0	0
AKAMAI TECHNOLOGIES INC	NOTE	00971TAL5	763	700,000	SH		DFND	15	700,000	0	0
AKEBIA THERAPEUTICS INC	COM	00972D105	597	176,230	SH		DFND	15	134,579	0	41,651
AKEBIA THERAPEUTICS INC	COM	00972D105	1,415	417,999	SH		DFND	48	417,999	0	0
AKEBIA THERAPEUTICS INC	COM	00972D105	7	2,000	SH		OTR	49	0	0	2,000
AKERNA CORP	COM	00973W102	8	1,699	SH		DFND	15	1,029	0	670
AKERNA CORP	COM	00973W102	301	60,950	SH		DFND	48	60,950	0	0
AKERO THERAPEUTICS INC	COM	00973Y108	554	19,106	SH		DFND	15	15,206	0	3,900
AKERO THERAPEUTICS INC	COM	00973Y108	1,031	35,542	SH		DFND	48	35,542	0	0
AKOUOS INC	COM	00973J101	251	18,103	SH		DFND	15	9,336	0	8,767
AKOUOS INC	COM	00973J101	706	50,925	SH		DFND	48	50,925	0	0
AKOUSTIS TECHNOLOGIES INC	COM	00973N102	278	20,850	SH		DFND	15	4,260	0	16,590
AKOUSTIS TECHNOLOGIES INC	COM	00973N102	1,301	97,501	SH		DFND	48	97,501	0	0
AKOUSTIS TECHNOLOGIES INC	COM	00973N102	2	165	SH		DFND	49	165	0	0
ALAMO GROUP INC	COM	011311107	8,729	55,902	SH		DFND	15	50,896	0	5,006
ALAMO GROUP INC	COM	011311107	3,997	25,595	SH		DFND	22	25,595	0	0
ALAMO GROUP INC	COM	011311107	153	979	SH		OTR	22,87	979	0	0
ALAMO GROUP INC	COM	011311107	16	100	SH		OTR	4	0	0	100
ALAMO GROUP INC	COM	011311107	3,899	24,969	SH		DFND	48	24,969	0	0
ALAMO GROUP INC	COM	011311107	62	396	SH		OTR	49	396	0	0
ALAMO GROUP INC	COM	011311107	67	427	SH		DFND	49	427	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	6,248	799,945	SH		OTR	15,66	799,945	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	17,652	2,260,153	SH		DFND	15,22	2,260,153	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	16,922	2,166,730	SH		DFND	15	1,774,032	0	392,698
ALAMOS GOLD INC-CLASS A	COM	011532108	26,202	3,354,980	SH		DFND	22	3,354,980	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	441	56,429	SH		DFND	42	56,429	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	2,796	358,058	SH		DFND	48	358,058	0	0
ALAMOS GOLD INC-CLASS A	COM	011532108	0	19	SH		DFND		19	0	0
ALARM.COM HOLDINGS INC	COM	011642105	18,336	212,275	SH		DFND	15	196,211	0	16,064
ALARM.COM HOLDINGS INC	COM	011642105	10,076	116,644	SH		DFND	22	116,644	0	0
ALARM.COM HOLDINGS INC	COM	011642105	3,620	41,913	SH		OTR	22,71	41,913	0	0
ALARM.COM HOLDINGS INC	COM	011642105	15	178	SH		DFND	4	178	0	0
ALARM.COM HOLDINGS INC	COM	011642105	10,242	118,573	SH		DFND	48	118,573	0	0
ALARM.COM HOLDINGS INC	COM	011642105	853	9,880	SH		DFND	49	9,880	0	0
ALARM.COM HOLDINGS INC	COM	011642105	809	9,367	SH		OTR	49	9,367	0	0
ALARM.COM HOLDINGS INC	COM	011642105	654	7,568	SH		DFND	59	7,301	0	267
ALARM.COM HOLDINGS INC	COM	011642105	0	1	SH		DFND		1	0	0
ALASKA AIR GROUP INC	COM	011659109	817	11,811	SH		DFND	15,22	11,811	0	0
ALASKA AIR GROUP INC	COM	011659109	300	4,330	SH		OTR	15,118	4,330	0	0
ALASKA AIR GROUP INC	COM	011659109	52,693	761,356	SH		DFND	15	568,465	0	192,891
ALASKA AIR GROUP INC	COM	011659109	148	2,144	SH		OTR	15,119	2,144	0	0
ALASKA AIR GROUP INC	COM	011659109	4,029	58,219	SH		DFND	22	58,219	0	0
ALASKA AIR GROUP INC	COM	011659109	4,518	65,277	SH		OTR	22,80	65,277	0	0
ALASKA AIR GROUP INC	COM	011659109	214	3,089	SH		DFND	4	3,089	0	0
ALASKA AIR GROUP INC	COM	011659109	26,542	383,498	SH		DFND	48	347,540	0	35,958
ALASKA AIR GROUP INC	COM	011659109	348	5,026	SH		OTR	49	4,968	58	0
ALASKA AIR GROUP INC	COM	011659109	3,033	43,826	SH		DFND	49	43,826	0	0
ALASKA AIR GROUP INC	COM	011659109	808	11,668	SH		DFND	59	11,668	0	0
ALASKA COMM SYSTEMS GROUP	COM	01167P101	125	38,344	SH		DFND	15	5,946	0	32,398
ALASKA COMM SYSTEMS GROUP	COM	01167P101	405	124,476	SH		DFND	48	124,476	0	0
ALASKA COMM SYSTEMS GROUP	COM	01167P101	31	9,600	SH		DFND	49	9,600	0	0
ALASKA COMM SYSTEMS GROUP	COM	01167P101	0	1	SH		DFND		1	0	0
ALBANY INTL CORP-CL A	CL A	012348108	15,737	188,539	SH		DFND	15	174,296	0	14,243
ALBANY INTL CORP-CL A	CL A	012348108	950	11,380	SH		OTR	22,83	11,380	0	0
ALBANY INTL CORP-CL A	CL A	012348108	1,103	13,215	SH		OTR	22,104	13,215	0	0
ALBANY INTL CORP-CL A	CL A	012348108	6,629	79,421	SH		DFND	22	79,421	0	0
ALBANY INTL CORP-CL A	CL A	012348108	8	90	SH		DFND	4	90	0	0
ALBANY INTL CORP-CL A	CL A	012348108	6,537	78,320	SH		DFND	48	78,320	0	0
ALBANY INTL CORP-CL A	CL A	012348108	60	717	SH		OTR	49	0	717	0
ALBANY INTL CORP-CL A	CL A	012348108	1,405	16,829	SH		DFND	49	16,829	0	0
ALBANY INTL CORP-CL A	CL A	012348108	1,457	17,451	SH		DFND	59	17,090	0	361
ALBEMARLE CORP	COM	012653101	582	3,986	SH		OTR	15,118	3,986	0	0
ALBEMARLE CORP	COM	012653101	369	2,524	SH		OTR	15,119	2,524	0	0
ALBEMARLE CORP	COM	012653101	1,395	9,548	SH		DFND	15,22	9,548	0	0
ALBEMARLE CORP	COM	012653101	69,539	475,939	SH		DFND	15	291,231	0	184,708
ALBEMARLE CORP	COM	012653101	4,856	33,234	SH		DFND	22	33,234	0	0
ALBEMARLE CORP	COM	012653101	54	372	SH		DFND	4	256	0	116
ALBEMARLE CORP	COM	012653101	1,271	8,699	SH		OTR	40,48	8,699	0	0
ALBEMARLE CORP	COM	012653101	225,272	1,541,794	SH		DFND	40	1,506,034	0	35,760
ALBEMARLE CORP	COM	012653101	1,307	8,947	SH		DFND	42	8,947	0	0
ALBEMARLE CORP	COM	012653101	53,064	363,177	SH		DFND	48	328,563	0	34,614
ALBEMARLE CORP	COM	012653101	650	4,452	SH		OTR	49	241	4,211	0
ALBEMARLE CORP	COM	012653101	3,956	27,078	SH		DFND	49	26,629	0	449
ALBEMARLE CORP	COM	012653101	52	356	SH		DFND	59	282	0	74
ALBEMARLE CORP	COM	012653101	20	140	SH		DFND		140	0	0
ALBERTSONS COS INC - CLASS A	COMM	013091103	19,600	1,027,773	SH		DFND	15	966,253	0	61,520
ALBERTSONS COS INC - CLASS A	COMM	013091103	3,612	189,407	SH		DFND	48	142,130	0	47,277
ALBERTSONS COS INC - CLASS A	COMM	013091103	15	795	SH		DFND	49	795	0	0
ALBIREO PHARMA INC	COM	01345P106	828	23,488	SH		DFND	15	14,305	0	9,183
ALBIREO PHARMA INC	COM	01345P106	702	19,910	SH		OTR	22,87	19,910	0	0
ALBIREO PHARMA INC	COM	01345P106	1,617	45,864	SH		DFND	48	45,864	0	0
ALBIREO PHARMA INC	COM	01345P106	129	3,660	SH		DFND	49	3,660	0	0
ALCOA CORP	COM	013872106	680	20,935	SH		OTR	15,119	20,935	0	0
ALCOA CORP	COM	013872106	9,759	300,373	SH		DFND	15	134,791	0	165,582
ALCOA CORP	COM	013872106	1,204	37,045	SH		OTR	22,69	37,045	0	0
ALCOA CORP	COM	013872106	11	333	SH		OTR	4	333	0	0
ALCOA CORP	COM	013872106	15,210	468,132	SH		DFND	48	468,132	0	0
ALCOA CORP	COM	013872106	261	8,039	SH		DFND	49	8,039	0	0
ALCOA CORP	COM	013872106	361	11,115	SH		OTR	49	9,505	456	1,154
ALCON INC	ORD	H01301128	297	4,225	SH		OTR	15,119	4,225	0	0
ALCON INC	ORD	H01301128	72,588	1,034,310	SH		DFND	15	872,747	0	161,563
ALCON INC	ORD	H01301128	32,976	469,882	SH		DFND	22	469,882	0	0
ALCON INC	ORD	H01301128	171	2,433	SH		DFND	4	2,433	0	0
ALCON INC	ORD	H01301128	126	1,789	SH		OTR	4	343	180	1,266
ALCON INC	ORD	H01301128	29,319	417,766	SH		DFND	40,22	417,766	0	0
ALCON INC	ORD	H01301128	11,917	169,811	SH		OTR	40,48	169,811	0	0
ALCON INC	ORD	H01301128	151,456	2,158,102	SH		DFND	40	2,156,121	0	1,981
ALCON INC	ORD	H01301128	4,467	63,650	SH		DFND	42	63,650	0	0
ALCON INC	ORD	H01301128	37,509	534,464	SH		DFND	48	534,464	0	0
ALCON INC	ORD	H01301128	3,514	50,071	SH		DFND	49	49,913	0	158
ALCON INC	ORD	H01301128	1,117	15,922	SH		OTR	49	8,800	4,835	2,287
ALCON INC	ORD	H01301128	538	7,659	SH		DFND	59	7,133	0	526
ALDEYRA THERAPEUTICS INC	COM	01438T106	285	23,950	SH		DFND	15	23,950	0	0
ALDEYRA THERAPEUTICS INC	COM	01438T106	278	23,404	SH		DFND	48	23,404	0	0
ALECTOR INC	COM	014442107	1,115	55,362	SH		DFND	15	41,371	0	13,991
ALECTOR INC	COM	014442107	2,557	126,949	SH		DFND	48	126,949	0	0
ALERIAN MLP ETF	ALER	00162Q452	523	17,147	SH		DFND	49	17,147	0	0
ALERIAN MLP ETF	ALER	00162Q452	1,967	64,477	SH		DFND	59	62,179	0	2,298
ALERUS FINANCIAL CORP	COM	01446U103	1,791	60,150	SH		DFND	15	50,850	0	9,300
ALERUS FINANCIAL CORP	COM	01446U103	1,079	36,225	SH		DFND	48	36,225	0	0
ALERUS FINANCIAL CORP	COM	01446U103	10	343	SH		OTR	49	0	343	0
ALERUS FINANCIAL CORP	COM	01446U103	0	1	SH		DFND		1	0	0
ALEXANDER & BALDWIN INC	COM	014491104	5,953	354,534	SH		DFND	15	288,452	0	66,082
ALEXANDER & BALDWIN INC	COM	014491104	3,165	188,491	SH		DFND	22	188,491	0	0
ALEXANDER & BALDWIN INC	COM	014491104	8	460	SH		DFND	4	460	0	0
ALEXANDER & BALDWIN INC	COM	014491104	2,917	173,724	SH		DFND	48	173,724	0	0
ALEXANDER & BALDWIN INC	COM	014491104	94	5,591	SH		OTR	49	5,591	0	0
ALEXANDER & BALDWIN INC	COM	014491104	1,183	70,475	SH		DFND	49	70,475	0	0
ALEXANDER & BALDWIN INC	COM	014491104	14	813	SH		DFND	59	813	0	0
ALEXANDER & BALDWIN INC	COM	014491104	1	47	SH		DFND		47	0	0
ALEXANDER'S INC	COM	014752109	5,844	21,075	SH		DFND	15	20,121	0	954
ALEXANDER'S INC	COM	014752109	1,776	6,403	SH		DFND	48	6,403	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	98,247	597,973	SH		DFND	15	358,248	0	239,725
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	1,896	11,541	SH		DFND	15,22	11,541	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	496	3,016	SH		OTR	15,119	3,016	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	716	4,358	SH		OTR	15,118	4,358	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	16,745	101,917	SH		DFND	22	101,917	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	109	665	SH		DFND	4	665	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	79,861	486,071	SH		DFND	48	445,281	0	40,790
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	3,048	18,549	SH		DFND	49	18,488	0	61
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	59	357	SH		OTR	49	250	107	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	222	1,349	SH		DFND	59	1,349	0	0
ALEXANDRIA REAL ESTATE EQUIT	COM	015271109	0	1	SH		DFND		1	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	1,133	7,411	SH		OTR	15,118	7,411	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	3,029	19,808	SH		DFND	15,22	19,808	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	146,474	957,913	SH		DFND	15	609,976	0	347,937
ALEXION PHARMACEUTICALS INC	COM	015351109	715	4,679	SH		OTR	15,119	4,679	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	8,864	57,968	SH		DFND	22	57,968	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	1,911	12,500	SH		OTR	4	0	12,500	0
ALEXION PHARMACEUTICALS INC	COM	015351109	177	1,157	SH		DFND	4	1,157	0	0
ALEXION PHARMACEUTICALS INC	COM	015351109	106,392	695,782	SH		DFND	48	631,210	0	64,572
ALEXION PHARMACEUTICALS INC	COM	015351109	643	4,202	SH		OTR	49	3,929	273	0
ALEXION PHARMACEUTICALS INC	COM	015351109	7,989	52,245	SH		DFND	49	52,058	0	187
ALEXION PHARMACEUTICALS INC	COM	015351109	0	1	SH		DFND		1	0	0
ALGONQUIN POWER & UTILITIES	COM	015857105	83	5,240	SH		OTR	15,119	5,240	0	0
ALGONQUIN POWER & UTILITIES	COM	015857105	9,868	623,004	SH		DFND	15	602,455	0	20,549
ALGONQUIN POWER & UTILITIES	COM	015857105	4,846	305,908	SH		DFND	48	305,908	0	0
ALGONQUIN POWER & UTILITIES	COM	015857105	0	1	SH		DFND		1	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	137,848	607,983	SH		DFND	15	569,402	0	38,581
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	7,796	34,386	SH		DFND	15,22	34,386	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	95,709	422,128	SH		DFND	22	422,128	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	119	525	SH		OTR	4	0	400	125
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	52	229	SH		DFND	4	62	0	167
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	13,678	60,326	SH		DFND	42	60,326	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	384,536	1,696,007	SH		DFND	48	1,696,007	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	3,211	14,162	SH		OTR	49	753	12,314	1,095
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	27,693	122,142	SH		DFND	49	121,522	0	620
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	10,076	44,440	SH		OTR	51,94	44,440	0	0
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	2,841	12,530	SH		DFND	51	6,230	0	6,300
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	2,852	12,580	SH		DFND	59	12,220	0	360
ALIBABA GROUP HOLDING-SP ADR	SPON	01609W102	760	3,350	SH		DFND		3,350	0	0
ALICO INC	COM	016230104	885	29,636	SH		DFND	15	26,395	0	3,241
ALICO INC	COM	016230104	403	13,494	SH		DFND	48	13,494	0	0
ALICO INC	COM	016230104	6	185	SH		DFND	49	185	0	0
ALIGN TECHNOLOGY INC	COM	016255101	186,172	343,788	SH		DFND	15	220,449	0	123,339
ALIGN TECHNOLOGY INC	COM	016255101	1,315	2,429	SH		OTR	15,118	2,429	0	0
ALIGN TECHNOLOGY INC	COM	016255101	88,914	164,191	SH		DFND	15,22	164,191	0	0
ALIGN TECHNOLOGY INC	COM	016255101	256	473	SH		OTR	15,119	473	0	0
ALIGN TECHNOLOGY INC	COM	016255101	9,546	17,628	SH		OTR	22,71	17,628	0	0
ALIGN TECHNOLOGY INC	COM	016255101	28,591	52,796	SH		DFND	22	52,796	0	0
ALIGN TECHNOLOGY INC	COM	016255101	325	601	SH		OTR	4	166	26	409
ALIGN TECHNOLOGY INC	COM	016255101	424	783	SH		DFND	4	783	0	0
ALIGN TECHNOLOGY INC	COM	016255101	2,417	4,463	SH		DFND	42	4,463	0	0
ALIGN TECHNOLOGY INC	COM	016255101	125,424	231,610	SH		DFND	48	207,431	0	24,179
ALIGN TECHNOLOGY INC	COM	016255101	15,472	28,570	SH		DFND	49	28,163	0	407
ALIGN TECHNOLOGY INC	COM	016255101	3,158	5,832	SH		OTR	49	5,044	788	0
ALIGN TECHNOLOGY INC	COM	016255101	663	1,224	SH		DFND	59	1,146	0	78
ALIGN TECHNOLOGY INC	COM	016255101	0	1	SH		DFND		1	0	0
ALIGOS THERAPEUTICS INC	COM	01626L105	80	3,504	SH		DFND	15	1,472	0	2,032
ALIGOS THERAPEUTICS INC	COM	01626L105	643	28,273	SH		DFND	48	28,273	0	0
ALKALINE WATER CO INC/THE	COM	01643A207	84	77,427	SH		DFND	48	77,427	0	0
ALKERMES PLC	SHS	G01767105	3,974	212,749	SH		OTR	15,66	212,749	0	0
ALKERMES PLC	SHS	G01767105	16,018	857,497	SH		DFND	15	637,997	0	219,500
ALKERMES PLC	SHS	G01767105	12,387	663,090	SH		DFND	22	663,090	0	0
ALKERMES PLC	SHS	G01767105	7,164	383,521	SH		DFND	48	338,233	0	45,288
ALKERMES PLC	SHS	G01767105	228	12,231	SH		OTR	49	12,231	0	0
ALKERMES PLC	SHS	G01767105	74	3,974	SH		DFND	49	3,974	0	0
ALKERMES PLC	SHS	G01767105	0	1	SH		DFND		1	0	0
ALLAKOS INC	COM	01671P100	3,858	33,609	SH		DFND	15	25,316	0	8,293
ALLAKOS INC	COM	01671P100	7,875	68,607	SH		DFND	48	68,607	0	0
ALLAKOS INC	COM	01671P100	10	90	SH		DFND	49	90	0	0
ALLEGHANY CORP	COM	017175100	35,119	56,074	SH		DFND	15	49,015	0	7,059
ALLEGHANY CORP	COM	017175100	485	775	SH		OTR	15,119	775	0	0
ALLEGHANY CORP	COM	017175100	9,378	14,974	SH		DFND	22	14,974	0	0
ALLEGHANY CORP	COM	017175100	5,010	7,999	SH		OTR	22,80	7,999	0	0
ALLEGHANY CORP	COM	017175100	103	164	SH		DFND	4	164	0	0
ALLEGHANY CORP	COM	017175100	33,343	53,239	SH		DFND	48	49,001	0	4,238
ALLEGHANY CORP	COM	017175100	27,457	43,840	SH		DFND	49	43,813	0	27
ALLEGHANY CORP	COM	017175100	62	99	SH		OTR	49	0	99	0
ALLEGHANY CORP	COM	017175100	115	183	SH		DFND	59	182	0	1
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	13,139	623,906	SH		DFND	15	508,442	0	115,464
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	150	7,146	SH		OTR	15,119	7,146	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	4,720	224,140	SH		OTR	22,96	224,140	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	6,910	328,105	SH		DFND	22	328,105	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	11	505	SH		DFND	4	505	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	6,804	323,094	SH		DFND	48	323,094	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	5	246	SH		OTR	49	10	236	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	302	14,353	SH		DFND	49	14,353	0	0
ALLEGHENY TECHNOLOGIES INC	COM	01741R102	0	1	SH		DFND		1	0	0
ALLEGIANCE BANCSHARES INC	COM	01748H107	4,198	103,561	SH		DFND	15	86,865	0	16,696
ALLEGIANCE BANCSHARES INC	COM	01748H107	1,903	46,943	SH		DFND	22	46,943	0	0
ALLEGIANCE BANCSHARES INC	COM	01748H107	1,951	48,129	SH		DFND	48	48,129	0	0
ALLEGIANCE BANCSHARES INC	COM	01748H107	319	7,874	SH		DFND	49	7,874	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	15,268	62,560	SH		DFND	15	51,622	0	10,938
ALLEGIANT TRAVEL CO	COM	01748X102	1,204	4,934	SH		OTR	22,69	4,934	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	3,685	15,100	SH		OTR	22,96	15,100	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	3,199	13,109	SH		OTR	22,71	13,109	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	8,260	33,845	SH		DFND	22	33,845	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	7,921	32,455	SH		DFND	48	32,455	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	1,068	4,378	SH		DFND	49	4,378	0	0
ALLEGIANT TRAVEL CO	COM	01748X102	0	1	SH		DFND		1	0	0
ALLEGION PLC	ORD	G0176J109	219	1,741	SH		OTR	15,119	1,741	0	0
ALLEGION PLC	ORD	G0176J109	393	3,132	SH		OTR	15,118	3,132	0	0
ALLEGION PLC	ORD	G0176J109	37,967	302,233	SH		DFND	15	155,871	0	146,362
ALLEGION PLC	ORD	G0176J109	1,037	8,254	SH		DFND	15,22	8,254	0	0
ALLEGION PLC	ORD	G0176J109	4,108	32,700	SH		DFND	22	32,700	0	0
ALLEGION PLC	ORD	G0176J109	54	429	SH		DFND	4	429	0	0
ALLEGION PLC	ORD	G0176J109	4	35	SH		OTR	4	0	0	35
ALLEGION PLC	ORD	G0176J109	36,632	291,609	SH		DFND	48	263,230	0	28,379
ALLEGION PLC	ORD	G0176J109	553	4,402	SH		OTR	49	2,157	2,245	0
ALLEGION PLC	ORD	G0176J109	2,573	20,481	SH		DFND	49	20,431	0	50
ALLEGION PLC	ORD	G0176J109	248	1,973	SH		DFND	59	1,916	0	57
ALLEGRO MICROSYSTEMS INC	COM	01749D105	791	31,187	SH		DFND	15	16,630	0	14,557
ALLEGRO MICROSYSTEMS INC	COM	01749D105	1,848	72,900	SH		DFND	48	58,380	0	14,520
ALLEGRO MICROSYSTEMS INC	COM	01749D105	3	130	SH		DFND	49	130	0	0
ALLENA PHARMACEUTICALS INC	COM	018119107	54	39,186	SH		DFND	48	39,186	0	0
ALLETE INC	COM	018522300	224	3,327	SH		OTR	15,119	3,327	0	0
ALLETE INC	COM	018522300	26,638	396,455	SH		DFND	15	350,267	0	46,188
ALLETE INC	COM	018522300	3,763	56,002	SH		DFND	22	56,002	0	0
ALLETE INC	COM	018522300	106	1,583	SH		OTR	22,87	1,583	0	0
ALLETE INC	COM	018522300	41	615	SH		DFND	4	615	0	0
ALLETE INC	COM	018522300	16,558	246,435	SH		DFND	48	246,435	0	0
ALLETE INC	COM	018522300	2,409	35,855	SH		DFND	49	35,725	0	130
ALLETE INC	COM	018522300	1,967	29,278	SH		OTR	49	29,203	75	0
ALLETE INC	COM	018522300	8	119	SH		DFND	59	119	0	0
ALLETE INC	COM	018522300	0	1	SH		DFND		1	0	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	51,514	459,576	SH		DFND	15	436,361	0	23,215
ALLIANCE DATA SYSTEMS CORP	COM	018581108	565	5,045	SH		OTR	15,119	5,045	0	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	402	3,590	SH		DFND	15,22	3,590	0	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	7,291	65,045	SH		DFND	22	65,045	0	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	92	821	SH		DFND	4	821	0	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	18	165	SH		OTR	4	0	78	87
ALLIANCE DATA SYSTEMS CORP	COM	018581108	20,455	182,488	SH		DFND	48	168,332	0	14,156
ALLIANCE DATA SYSTEMS CORP	COM	018581108	4,158	37,094	SH		DFND	49	36,936	0	158
ALLIANCE DATA SYSTEMS CORP	COM	018581108	77	691	SH		OTR	49	455	236	0
ALLIANCE DATA SYSTEMS CORP	COM	018581108	1	5	SH		DFND	59	5	0	0
ALLIANT ENERGY CORP	COM	018802108	1,213	22,394	SH		DFND	15,22	22,394	0	0
ALLIANT ENERGY CORP	COM	018802108	586	10,811	SH		OTR	15,119	10,811	0	0
ALLIANT ENERGY CORP	COM	018802108	49,743	918,448	SH		DFND	15	518,011	0	400,437
ALLIANT ENERGY CORP	COM	018802108	467	8,630	SH		OTR	15,118	8,630	0	0
ALLIANT ENERGY CORP	COM	018802108	2,253	41,605	SH		DFND	22	41,605	0	0
ALLIANT ENERGY CORP	COM	018802108	3	47	SH		OTR	4	0	0	47
ALLIANT ENERGY CORP	COM	018802108	37	682	SH		DFND	4	682	0	0
ALLIANT ENERGY CORP	COM	018802108	51,456	950,071	SH		DFND	48	875,053	0	75,018
ALLIANT ENERGY CORP	COM	018802108	241	4,446	SH		OTR	49	1,732	2,714	0
ALLIANT ENERGY CORP	COM	018802108	5,128	94,677	SH		DFND	49	94,677	0	0
ALLIANT ENERGY CORP	COM	018802108	0	1	SH		DFND	59	1	0	0
ALLIED ESPORTS ENTERTAINMENT	COM	019170109	58	20,241	SH		DFND	48	20,241	0	0
ALLIED MOTION TECHNOLOGIES	COM	019330109	1,160	22,604	SH		DFND	15	21,384	0	1,220
ALLIED MOTION TECHNOLOGIES	COM	019330109	1,293	25,198	SH		OTR	22,83	25,198	0	0
ALLIED MOTION TECHNOLOGIES	COM	019330109	1,018	19,829	SH		DFND	48	19,829	0	0
ALLISON TRANSMISSION HOLDING	COM	01973R101	220	5,398	SH		OTR	15,119	5,398	0	0
ALLISON TRANSMISSION HOLDING	COM	01973R101	15,766	386,141	SH		DFND	15	329,853	0	56,288
ALLISON TRANSMISSION HOLDING	COM	01973R101	3,680	90,134	SH		OTR	22,80	90,134	0	0
ALLISON TRANSMISSION HOLDING	COM	01973R101	2	39	SH		OTR	4	0	0	39
ALLISON TRANSMISSION HOLDING	COM	01973R101	8,650	211,844	SH		DFND	48	176,896	0	34,948
ALLISON TRANSMISSION HOLDING	COM	01973R101	2	56	SH		OTR	49	0	56	0
ALLISON TRANSMISSION HOLDING	COM	01973R101	275	6,740	SH		DFND	49	6,740	0	0
ALLISON TRANSMISSION HOLDING	COM	01973R101	1,811	44,357	SH		DFND	59	42,782	0	1,575
ALLOGENE THERAPEUTICS INC	COM	019770106	2,708	76,725	SH		DFND	15	56,828	0	19,897
ALLOGENE THERAPEUTICS INC	COM	019770106	5,133	145,424	SH		DFND	48	145,424	0	0
ALLOGENE THERAPEUTICS INC	COM	019770106	0	1	SH		DFND		1	0	0
ALLOVIR INC	COM	019818103	541	23,119	SH		DFND	15	13,954	0	9,165
ALLOVIR INC	COM	019818103	1,428	61,042	SH		DFND	48	61,042	0	0
ALLSCRIPTS HEALTHCARE SOLUTI	COM	01988P108	11,500	765,916	SH		DFND	15	630,857	0	135,059
ALLSCRIPTS HEALTHCARE SOLUTI	COM	01988P108	5,476	364,721	SH		DFND	22	364,721	0	0
ALLSCRIPTS HEALTHCARE SOLUTI	COM	01988P108	64	4,240	SH		DFND	4	4,240	0	0
ALLSCRIPTS HEALTHCARE SOLUTI	COM	01988P108	5,927	394,713	SH		DFND	48	394,713	0	0
ALLSCRIPTS HEALTHCARE SOLUTI	COM	01988P108	931	61,998	SH		DFND	49	61,998	0	0
ALLSTATE CORP	COM	020002101	1,173	10,210	SH		OTR	15,118	10,210	0	0
ALLSTATE CORP	COM	020002101	202,696	1,764,107	SH		DFND	15	1,250,351	0	513,756
ALLSTATE CORP	COM	020002101	3,125	27,194	SH		DFND	15,22	27,194	0	0
ALLSTATE CORP	COM	020002101	2,465	21,451	SH		OTR	15,119	21,451	0	0
ALLSTATE CORP	COM	020002101	6,349	55,259	SH		DFND	22	55,259	0	0
ALLSTATE CORP	COM	020002101	330	2,871	SH		DFND	4	2,871	0	0
ALLSTATE CORP	COM	020002101	126	1,093	SH		OTR	4	1,058	0	35
ALLSTATE CORP	COM	020002101	106,134	923,707	SH		DFND	48	832,067	0	91,640
ALLSTATE CORP	COM	020002101	3,409	29,666	SH		OTR	49	16,812	12,228	626
ALLSTATE CORP	COM	020002101	18,670	162,489	SH		DFND	49	160,324	0	2,165
ALLSTATE CORP	COM	020002101	5,524	48,076	SH		DFND	59	47,128	0	948
ALLSTATE CORP	COM	020002101	118	1,027	SH		DFND		1,027	0	0
ALLY FINANCIAL INC	COM	02005N100	103,991	2,300,185	SH		DFND	15	2,008,832	0	291,353
ALLY FINANCIAL INC	COM	02005N100	7,249	160,342	SH		OTR	15,118	160,342	0	0
ALLY FINANCIAL INC	COM	02005N100	1,269	28,078	SH		OTR	15,119	28,078	0	0
ALLY FINANCIAL INC	COM	02005N100	35,084	776,023	SH		DFND	22	776,023	0	0
ALLY FINANCIAL INC	COM	02005N100	565	12,495	SH		DFND	4	12,495	0	0
ALLY FINANCIAL INC	COM	02005N100	4	90	SH		OTR	4	90	0	0
ALLY FINANCIAL INC	COM	02005N100	149	3,291	SH		DFND	42	3,291	0	0
ALLY FINANCIAL INC	COM	02005N100	36,272	802,298	SH		DFND	48	689,549	0	112,749
ALLY FINANCIAL INC	COM	02005N100	427	9,442	SH		OTR	49	6,043	3,329	70
ALLY FINANCIAL INC	COM	02005N100	6,672	147,567	SH		DFND	49	144,575	0	2,992
ALLY FINANCIAL INC	COM	02005N100	482	10,657	SH		DFND	59	9,330	0	1,327
ALLY FINANCIAL INC	COM	02005N100	0	1	SH		DFND		1	0	0
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	29,197	206,793	SH		DFND	15	148,477	0	58,316
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	1,815	12,857	SH		DFND	22	12,857	0	0
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	4	26	SH		DFND	4	26	0	0
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	26,355	186,663	SH		DFND	48	150,990	0	35,673
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	1,873	13,264	SH		DFND	49	13,264	0	0
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	291	2,061	SH		OTR	49	2,061	0	0
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	21	149	SH		DFND	59	119	0	30
ALNYLAM PHARMACEUTICALS INC	COM	02043Q107	0	1	SH		DFND		1	0	0
ALPHA & OMEGA SEMICONDUCTOR	SHS	G6331P104	886	27,101	SH		DFND	15	5,213	0	21,888
ALPHA & OMEGA SEMICONDUCTOR	SHS	G6331P104	1,789	54,713	SH		DFND	48	54,713	0	0
ALPHA METALLURGICAL RESOURCE	COM	020764106	613	48,569	SH		DFND	48	48,569	0	0
ALPHA METALLURGICAL RESOURCE	COM	020764106	0	3	SH		DFND		3	0	0
ALPHA PRO TECH LTD	COM	020772109	296	30,278	SH		DFND	15	29,372	0	906
ALPHA PRO TECH LTD	COM	020772109	272	27,870	SH		DFND	48	27,870	0	0
ALPHA PRO TECH LTD	COM	020772109	7	735	SH		DFND	49	735	0	0
ALPHA PRO TECH LTD	COM	020772109	160	16,424	SH		DFND	59	16,424	0	0
ALPHABET INC-CL A	CAP	02079K305	55,742	27,026	SH		DFND	15,22	27,026	0	0
ALPHABET INC-CL A	CAP	02079K305	37,443	18,154	SH		OTR	15,118	18,154	0	0
ALPHABET INC-CL A	CAP	02079K305	8,372	4,059	SH		OTR	15,119	4,059	0	0
ALPHABET INC-CL A	CAP	02079K305	2,504,334	1,214,211	SH		DFND	15	696,553	0	517,658
ALPHABET INC-CL A	CAP	02079K305	232,122	112,543	SH		DFND	22	112,543	0	0
ALPHABET INC-CL A	CAP	02079K305	18,160	8,805	SH		DFND	4	8,805	0	0
ALPHABET INC-CL A	CAP	02079K305	5,129	2,487	SH		OTR	4	371	2,112	4
ALPHABET INC-CL A	CAP	02079K305	731,101	354,470	SH		DFND	40	319,313	0	35,157
ALPHABET INC-CL A	CAP	02079K305	18,897	9,162	SH		OTR	40,48	9,162	0	0
ALPHABET INC-CL A	CAP	02079K305	29,463	14,285	SH		DFND	40,22	14,285	0	0
ALPHABET INC-CL A	CAP	02079K305	7,508	3,640	SH		DFND	42	3,640	0	0
ALPHABET INC-CL A	CAP	02079K305	2,006,269	972,727	SH		DFND	48	880,066	0	92,661
ALPHABET INC-CL A	CAP	02079K305	143,063	69,363	SH		OTR	49	20,035	18,815	30,513
ALPHABET INC-CL A	CAP	02079K305	740,562	359,057	SH		DFND	49	354,213	86	4,758
ALPHABET INC-CL A	CAP	02079K305	118,240	57,328	SH		DFND	51	24,628	0	32,700
ALPHABET INC-CL A	CAP	02079K305	13,669	6,627	SH		DFND	59	6,427	0	200
ALPHABET INC-CL A	CAP	02079K305	919	445	SH		DFND		445	0	0
ALPHABET INC-CL C	CAP	02079K107	20,155	9,743	SH		OTR	15,118	9,743	0	0
ALPHABET INC-CL C	CAP	02079K107	2,301,289	1,112,470	SH		DFND	15	622,733	0	489,737
ALPHABET INC-CL C	CAP	02079K107	53,526	25,875	SH		DFND	15,22	25,875	0	0
ALPHABET INC-CL C	CAP	02079K107	8,101	3,916	SH		OTR	15,119	3,916	0	0
ALPHABET INC-CL C	CAP	02079K107	401,939	194,302	SH		DFND	22	194,302	0	0
ALPHABET INC-CL C	CAP	02079K107	9,373	4,531	SH		DFND	4	4,502	0	29
ALPHABET INC-CL C	CAP	02079K107	8,984	4,343	SH		OTR	4	463	3,880	0
ALPHABET INC-CL C	CAP	02079K107	962	465	SH		DFND	40	465	0	0
ALPHABET INC-CL C	CAP	02079K107	443	214	SH		DFND	42	214	0	0
ALPHABET INC-CL C	CAP	02079K107	1,804,464	872,299	SH		DFND	48	782,547	0	89,752
ALPHABET INC-CL C	CAP	02079K107	333,141	161,044	SH		DFND	49	158,700	15	2,329
ALPHABET INC-CL C	CAP	02079K107	42,641	20,613	SH		OTR	49	8,630	11,772	211
ALPHABET INC-CL C	CAP	02079K107	1,424,277	688,512	SH		DFND	51	491,073	0	197,439
ALPHABET INC-CL C	CAP	02079K107	29,838	14,424	SH		OTR	51,48	14,424	0	0
ALPHABET INC-CL C	CAP	02079K107	61,238	29,603	SH		DFND	51,22	29,603	0	0
ALPHABET INC-CL C	CAP	02079K107	217,773	105,274	SH		OTR	51,94	105,274	0	0
ALPHABET INC-CL C	CAP	02079K107	10,758	5,201	SH		DFND	59	4,942	0	259
ALPHABET INC-CL C	CAP	02079K107	250	121	SH		DFND		121	0	0
ALPHATEC HOLDINGS INC	COM	02081G201	1,161	73,527	SH		DFND	15	48,096	0	25,431
ALPHATEC HOLDINGS INC	COM	02081G201	1,273	80,624	SH		OTR	22,69	80,624	0	0
ALPHATEC HOLDINGS INC	COM	02081G201	2,351	148,891	SH		DFND	48	148,891	0	0
ALPHATEC HOLDINGS INC	COM	02081G201	3	209	SH		DFND	49	209	0	0
ALPINE IMMUNE SCIENCES INC	COM	02083G100	163	15,380	SH		DFND	15	15,380	0	0
ALPINE IMMUNE SCIENCES INC	COM	02083G100	77	7,304	SH		DFND	48	7,304	0	0
ALPINE INCOME PROPERTY TRUST	COM	02083X103	568	32,724	SH		DFND	15	28,761	0	3,963
ALPINE INCOME PROPERTY TRUST	COM	02083X103	285	16,424	SH		DFND	48	16,424	0	0
ALPINE INCOME PROPERTY TRUST	COM	02083X103	3	190	SH		DFND	49	190	0	0
ALTA EQUIPMENT GROUP INC	COMM	02128L106	156	11,999	SH		DFND	15	1,952	0	10,047
ALTA EQUIPMENT GROUP INC	COMM	02128L106	540	41,531	SH		DFND	48	41,531	0	0
ALTA EQUIPMENT GROUP INC	COMM	02128L106	7	509	SH		DFND	49	509	0	0
ALTABANCORP	COM	021347109	2,736	65,069	SH		DFND	15	52,488	0	12,581
ALTABANCORP	COM	021347109	1,705	40,562	SH		DFND	48	40,562	0	0
ALTAIR ENGINEERING INC	NOTE	021369AA1	290	200,000	SH		DFND	15	200,000	0	0
ALTAIR ENGINEERING INC - A	COM	021369103	2,671	42,691	SH		DFND	15	26,687	0	16,004
ALTAIR ENGINEERING INC - A	COM	021369103	6,855	109,561	SH		DFND	48	109,561	0	0
ALTAIR ENGINEERING INC - A	COM	021369103	119	1,900	SH		OTR	49	0	0	1,900
ALTAIR ENGINEERING INC - A	COM	021369103	15	243	SH		DFND	59	243	0	0
ALTERYX INC	NOTE	02156BAD5	281	300,000	SH		DFND	15	300,000	0	0
ALTERYX INC	NOTE	02156BAF0	271	300,000	SH		DFND	15	300,000	0	0
ALTERYX INC - CLASS A	COM	02156B103	5,975	72,025	SH		DFND	15	44,895	0	27,130
ALTERYX INC - CLASS A	COM	02156B103	41	500	SH		DFND	4	500	0	0
ALTERYX INC - CLASS A	COM	02156B103	7,699	92,800	SH		DFND	48	76,221	0	16,579
ALTERYX INC - CLASS A	COM	02156B103	311	3,752	SH		DFND	49	3,752	0	0
ALTICE USA INC- A	CL A	02156K103	14,403	442,774	SH		DFND	15	325,280	0	117,494
ALTICE USA INC- A	CL A	02156K103	492	15,129	SH		OTR	15,119	15,129	0	0
ALTICE USA INC- A	CL A	02156K103	2,784	85,570	SH		OTR	22,80	85,570	0	0
ALTICE USA INC- A	CL A	02156K103	651	20,010	SH		DFND	22	20,010	0	0
ALTICE USA INC- A	CL A	02156K103	12,888	396,203	SH		DFND	48	323,208	0	72,995
ALTICE USA INC- A	CL A	02156K103	49	1,492	SH		DFND	49	1,492	0	0
ALTICE USA INC- A	CL A	02156K103	5	140	SH		OTR	49	140	0	0
ALTIMMUNE INC	COM	02155H200	782	55,357	SH		DFND	48	55,357	0	0
ALTISOURCE PORTFOLIO SOL	REG	L0175J104	90	9,791	SH		DFND	15	1,956	0	7,835
ALTISOURCE PORTFOLIO SOL	REG	L0175J104	157	17,062	SH		DFND	48	17,062	0	0
ALTO INGREDIENTS INC	COM	021513106	677	124,594	SH		DFND	15	124,594	0	0
ALTO INGREDIENTS INC	COM	021513106	171	31,532	SH		DFND	48	31,532	0	0
ALTO INGREDIENTS INC	COM	021513106	0	25	SH		DFND		25	0	0
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	7,366	133,145	SH		DFND	15	75,677	0	57,468
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	1,404	25,375	SH		OTR	22,104	25,375	0	0
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	2,904	52,502	SH		OTR	22,80	52,502	0	0
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	8,704	157,343	SH		DFND	48	157,343	0	0
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	72	1,301	SH		DFND	49	1,301	0	0
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	1,178	21,291	SH		DFND	59	20,783	0	508
ALTRA INDUSTRIAL MOTION CORP	COM	02208R106	2	34	SH		DFND		34	0	0
ALTRIA GROUP INC	COM	02209S103	8,506	166,272	SH		DFND	15,22	166,272	0	0
ALTRIA GROUP INC	COM	02209S103	5,341	104,395	SH		OTR	15,119	104,395	0	0
ALTRIA GROUP INC	COM	02209S103	689,687	13,480,977	SH		DFND	15	12,245,267	0	1,235,710
ALTRIA GROUP INC	COM	02209S103	3,211	62,772	SH		OTR	15,118	62,772	0	0
ALTRIA GROUP INC	COM	02209S103	46,680	912,433	SH		DFND	22	912,433	0	0
ALTRIA GROUP INC	COM	02209S103	61	1,200	SH		OTR	4	150	1,050	0
ALTRIA GROUP INC	COM	02209S103	1,658	32,406	SH		DFND	4	32,406	0	0
ALTRIA GROUP INC	COM	02209S103	1,886	36,872	SH		OTR	40,48	36,872	0	0
ALTRIA GROUP INC	COM	02209S103	69,524	1,358,950	SH		DFND	40	1,206,537	0	152,413
ALTRIA GROUP INC	COM	02209S103	1,865	36,453	SH		DFND	42	36,453	0	0
ALTRIA GROUP INC	COM	02209S103	281,996	5,512,043	SH		DFND	48	4,943,061	0	568,982
ALTRIA GROUP INC	COM	02209S103	36,699	717,332	SH		DFND	49	714,993	0	2,339
ALTRIA GROUP INC	COM	02209S103	6,047	118,191	SH		OTR	49	67,659	50,332	200
ALTRIA GROUP INC	COM	02209S103	4,479	87,555	SH		DFND	59	85,752	0	1,802
ALTRIA GROUP INC	COM	02209S103	0	1	SH		DFND		1	0	0
ALX ONCOLOGY HOLDINGS INC	COM	00166B105	1,122	15,210	SH		DFND	15	8,719	0	6,491
ALX ONCOLOGY HOLDINGS INC	COM	00166B105	891	12,089	SH		OTR	22,78	12,089	0	0
ALX ONCOLOGY HOLDINGS INC	COM	00166B105	2,800	37,973	SH		DFND	48	37,973	0	0
AMALGAMATED FINANCIAL CORP	COM	022671101	1,370	82,605	SH		DFND	15	73,987	0	8,618
AMALGAMATED FINANCIAL CORP	COM	022671101	665	40,084	SH		DFND	48	40,084	0	0
A-MARK PRECIOUS METALS INC	COM	00181T107	1,015	28,203	SH		DFND	15	25,206	0	2,997
A-MARK PRECIOUS METALS INC	COM	00181T107	411	11,428	SH		DFND	48	11,428	0	0
A-MARK PRECIOUS METALS INC	COM	00181T107	10	270	SH		DFND	49	270	0	0
AMAZON.COM INC	COM	023135106	44,771	14,470	SH		OTR	15,118	14,470	0	0
AMAZON.COM INC	COM	023135106	118,667	38,353	SH		DFND	15,22	38,353	0	0
AMAZON.COM INC	COM	023135106	11,986	3,874	SH		OTR	15,119	3,874	0	0
AMAZON.COM INC	COM	023135106	5,518,808	1,783,667	SH		DFND	15	1,062,402	0	721,265
AMAZON.COM INC	COM	023135106	632,136	204,305	SH		DFND	22	204,305	0	0
AMAZON.COM INC	COM	023135106	9,347	3,021	SH		OTR	4	357	2,607	57
AMAZON.COM INC	COM	023135106	26,423	8,540	SH		DFND	4	8,522	0	18
AMAZON.COM INC	COM	023135106	407,125	131,582	SH		DFND	40	123,692	0	7,890
AMAZON.COM INC	COM	023135106	16,454	5,318	SH		OTR	40,48	5,318	0	0
AMAZON.COM INC	COM	023135106	25,721	8,313	SH		DFND	40,22	8,313	0	0
AMAZON.COM INC	COM	023135106	6,810	2,201	SH		DFND	42	2,201	0	0
AMAZON.COM INC	COM	023135106	4,107,484	1,327,530	SH		DFND	48	1,195,948	0	131,582
AMAZON.COM INC	COM	023135106	1,148,126	371,072	SH		DFND	49	364,325	76	6,671
AMAZON.COM INC	COM	023135106	123,336	39,862	SH		OTR	49	24,720	14,948	194
AMAZON.COM INC	COM	023135106	28,650	9,260	SH		DFND	59	9,030	0	230
AMAZON.COM INC	COM	023135106	558	180	SH		DFND		180	0	0
AMBAC FINANCIAL GROUP INC	COM	023139884	3,560	212,665	SH		DFND	15	167,649	0	45,016
AMBAC FINANCIAL GROUP INC	COM	023139884	2,020	120,681	SH		DFND	22	120,681	0	0
AMBAC FINANCIAL GROUP INC	COM	023139884	2	125	SH		DFND	4	125	0	0
AMBAC FINANCIAL GROUP INC	COM	023139884	1,916	114,449	SH		DFND	48	114,449	0	0
AMBAC FINANCIAL GROUP INC	COM	023139884	231	13,776	SH		DFND	49	13,776	0	0
AMBARELLA INC	SHS	G037AX101	5,062	50,428	SH		DFND	15	26,711	0	23,717
AMBARELLA INC	SHS	G037AX101	956	9,518	SH		OTR	22,69	9,518	0	0
AMBARELLA INC	SHS	G037AX101	8,577	85,435	SH		DFND	48	85,435	0	0
AMBARELLA INC	SHS	G037AX101	110	1,095	SH		OTR	49	0	1,095	0
AMBEV SA-ADR	SPON	02319V103	19	6,879	SH		OTR	4	0	0	6,879
AMBEV SA-ADR	SPON	02319V103	4,653	1,698,014	SH		DFND	40	1,698,014	0	0
AMBEV SA-ADR	SPON	02319V103	73	26,681	SH		OTR	49	1,318	560	24,803
AMBEV SA-ADR	SPON	02319V103	3,972	1,449,600	SH		OTR	51,94	1,449,600	0	0
AMBEV SA-ADR	SPON	02319V103	1,217	444,300	SH		DFND	51	217,900	0	226,400
AMC ENTERTAINMENT HLDS-CL A	CL A	00165C104	6,145	601,880	SH		DFND	15	279,436	0	322,444
AMC ENTERTAINMENT HLDS-CL A	CL A	00165C104	8,713	853,400	SH		DFND	48	853,400	0	0
AMC ENTERTAINMENT HLDS-CL A	CL A	00165C104	44	4,317	SH		DFND	49	4,317	0	0
AMC ENTERTAINMENT HLDS-CL A	CL A	00165C104	0	1	SH		DFND		1	0	0
AMC NETWORKS INC-A	CL A	00164V103	12,102	227,649	SH		DFND	15	209,445	0	18,204
AMC NETWORKS INC-A	CL A	00164V103	149	2,811	SH		OTR	15,119	2,811	0	0
AMC NETWORKS INC-A	CL A	00164V103	4,152	78,113	SH		DFND	22	78,113	0	0
AMC NETWORKS INC-A	CL A	00164V103	3	55	SH		DFND	4	55	0	0
AMC NETWORKS INC-A	CL A	00164V103	3,985	74,966	SH		DFND	48	74,966	0	0
AMC NETWORKS INC-A	CL A	00164V103	566	10,639	SH		DFND	49	10,639	0	0
AMC NETWORKS INC-A	CL A	00164V103	385	7,251	SH		DFND	59	7,251	0	0
AMC NETWORKS INC-A	CL A	00164V103	0	1	SH		DFND		1	0	0
AMCOR PLC	ORD	G0250X107	613	52,517	SH		OTR	15,118	52,517	0	0
AMCOR PLC	ORD	G0250X107	1,643	140,644	SH		DFND	15,22	140,644	0	0
AMCOR PLC	ORD	G0250X107	53	4,532	SH		OTR	15,119	4,532	0	0
AMCOR PLC	ORD	G0250X107	88,906	7,611,851	SH		DFND	15	5,120,102	0	2,491,749
AMCOR PLC	ORD	G0250X107	3,643	311,914	SH		DFND	22	311,914	0	0
AMCOR PLC	ORD	G0250X107	69	5,865	SH		DFND	4	5,865	0	0
AMCOR PLC	ORD	G0250X107	53,337	4,566,542	SH		DFND	48	4,092,195	0	474,347
AMCOR PLC	ORD	G0250X107	3,330	285,115	SH		DFND	49	284,145	0	970
AMCOR PLC	ORD	G0250X107	22	1,882	SH		OTR	49	1,822	60	0
AMCOR PLC	ORD	G0250X107	14	1,170	SH		DFND	59	1,124	0	46
AMCOR PLC	ORD	G0250X107	0	1	SH		DFND		1	0	0
AMDOCS LTD	SHS	G02602103	14,334	204,328	SH		DFND	15	143,188	0	61,140
AMDOCS LTD	SHS	G02602103	278	3,963	SH		OTR	15,119	3,963	0	0
AMDOCS LTD	SHS	G02602103	1,062	15,141	SH		DFND	22	15,141	0	0
AMDOCS LTD	SHS	G02602103	1,991	28,384	SH		OTR	22,80	28,384	0	0
AMDOCS LTD	SHS	G02602103	11	150	SH		OTR	4	150	0	0
AMDOCS LTD	SHS	G02602103	37	530	SH		DFND	4	530	0	0
AMDOCS LTD	SHS	G02602103	10,985	156,593	SH		DFND	48	117,299	0	39,294
AMDOCS LTD	SHS	G02602103	649	9,248	SH		OTR	49	6,790	2,458	0
AMDOCS LTD	SHS	G02602103	2,802	39,942	SH		DFND	49	39,942	0	0
AMDOCS LTD	SHS	G02602103	1,849	26,355	SH		DFND	59	25,533	0	822
AMDOCS LTD	SHS	G02602103	0	1	SH		DFND		1	0	0
AMEDISYS INC	COM	023436108	69	260	SH		OTR	15,119	260	0	0
AMEDISYS INC	COM	023436108	44,016	166,228	SH		DFND	15	151,740	0	14,488
AMEDISYS INC	COM	023436108	474	1,790	SH		DFND	15,22	1,790	0	0
AMEDISYS INC	COM	023436108	12,105	45,716	SH		DFND	22	45,716	0	0
AMEDISYS INC	COM	023436108	155	586	SH		DFND	4	586	0	0
AMEDISYS INC	COM	023436108	32,693	123,469	SH		DFND	48	113,532	0	9,937
AMEDISYS INC	COM	023436108	1,121	4,232	SH		OTR	49	4,057	175	0
AMEDISYS INC	COM	023436108	4,894	18,483	SH		DFND	49	18,418	0	65
AMEDISYS INC	COM	023436108	1,492	5,634	SH		DFND	59	5,475	0	159
AMEDISYS INC	COM	023436108	0	1	SH		DFND		1	0	0
AMER NATL BNKSHS/DANVILLE VA	COM	027745108	1,468	44,403	SH		DFND	15	35,323	0	9,080
AMER NATL BNKSHS/DANVILLE VA	COM	027745108	998	30,174	SH		DFND	48	30,174	0	0
AMER NATL BNKSHS/DANVILLE VA	COM	027745108	0	1	SH		DFND		1	0	0
AMERANT BANCORP INC	CL A	023576101	605	32,559	SH		DFND	15	13,326	0	19,233
AMERANT BANCORP INC	CL A	023576101	970	52,244	SH		DFND	48	52,244	0	0
AMERCO	COM	023586100	179	293	SH		OTR	15,119	293	0	0
AMERCO	COM	023586100	11,526	18,815	SH		DFND	15	13,919	0	4,896
AMERCO	COM	023586100	9,676	15,795	SH		DFND	48	13,077	0	2,718
AMERCO	COM	023586100	9	14	SH		OTR	49	4	10	0
AMERCO	COM	023586100	272	444	SH		DFND	49	444	0	0
AMEREN CORPORATION	COM	023608102	701	8,622	SH		OTR	15,118	8,622	0	0
AMEREN CORPORATION	COM	023608102	1,002	12,310	SH		OTR	15,119	12,310	0	0
AMEREN CORPORATION	COM	023608102	73,894	908,238	SH		DFND	15	511,295	0	396,943
AMEREN CORPORATION	COM	023608102	1,807	22,213	SH		DFND	15,22	22,213	0	0
AMEREN CORPORATION	COM	023608102	7,752	95,276	SH		DFND	22	95,276	0	0
AMEREN CORPORATION	COM	023608102	2	26	SH		OTR	4	0	0	26
AMEREN CORPORATION	COM	023608102	157	1,935	SH		DFND	4	1,935	0	0
AMEREN CORPORATION	COM	023608102	55,397	680,889	SH		DFND	48	606,970	0	73,919
AMEREN CORPORATION	COM	023608102	335	4,122	SH		OTR	49	3,165	957	0
AMEREN CORPORATION	COM	023608102	5,765	70,858	SH		DFND	49	70,657	0	201
AMEREN CORPORATION	COM	023608102	1,541	18,936	SH		DFND	59	18,296	0	640
AMERESCO INC-CL A	CL A	02361E108	1,650	33,924	SH		DFND	15	26,316	0	7,608
AMERESCO INC-CL A	CL A	02361E108	1,516	31,180	SH		OTR	22,83	31,180	0	0
AMERESCO INC-CL A	CL A	02361E108	2,448	50,331	SH		OTR	22,78	50,331	0	0
AMERESCO INC-CL A	CL A	02361E108	2,911	59,853	SH		DFND	48	59,853	0	0
AMERESCO INC-CL A	CL A	02361E108	10	210	SH		DFND	49	210	0	0
AMERESCO INC-CL A	CL A	02361E108	97	2,000	SH		OTR	49	0	2,000	0
AMERESCO INC-CL A	CL A	02361E108	118	2,422	SH		DFND	59	2,320	0	102
AMERICA MOVIL-SPN ADR CL L	SPON	02364W105	6,561	483,142	SH		DFND	22	483,142	0	0
AMERICA MOVIL-SPN ADR CL L	SPON	02364W105	599	44,073	SH		DFND	49	44,073	0	0
AMERICA MOVIL-SPN ADR CL L	SPON	02364W105	43	3,155	SH		OTR	49	1,200	1,955	0
AMERICA MOVIL-SPN ADR CL L	SPON	02364W105	54	4,012	SH		DFND	59	4,012	0	0
AMERICA MOVIL-SPN ADR CL L	SPON	02364W105	0	1	SH		DFND		1	0	0
AMERICAN AIRLINES GROUP	NOTE	02376RAF9	1,202	700,000	SH		DFND	15	700,000	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	474	19,830	SH		OTR	15,119	19,830	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	42,687	1,786,047	SH		DFND	15	814,724	0	971,323
AMERICAN AIRLINES GROUP INC	COM	02376R102	519	21,725	SH		OTR	15,118	21,725	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	1,387	58,039	SH		DFND	15,22	58,039	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	3,059	128,010	SH		DFND	22	128,010	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	45	1,898	SH		DFND	4	1,898	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	45,445	1,901,448	SH		DFND	48	1,714,781	0	186,667
AMERICAN AIRLINES GROUP INC	COM	02376R102	30	1,273	SH		OTR	49	738	535	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	2,165	90,565	SH		DFND	49	86,799	0	3,766
AMERICAN AIRLINES GROUP INC	COM	02376R102	0	1	SH		DFND	59	1	0	0
AMERICAN AIRLINES GROUP INC	COM	02376R102	0	11	SH		DFND		11	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	11,185	344,790	SH		DFND	15	300,639	0	44,151
AMERICAN ASSETS TRUST INC	COM	024013104	4,218	130,022	SH		DFND	22	130,022	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	1,012	31,203	SH		OTR	22,86	31,203	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	59	1,830	SH		DFND	4	1,830	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	5,512	169,908	SH		DFND	48	169,908	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	1,580	48,695	SH		DFND	49	48,695	0	0
AMERICAN ASSETS TRUST INC	COM	024013104	13	399	SH		DFND	59	399	0	0
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	6,267	648,761	SH		DFND	15	542,905	0	105,856
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	231	23,930	SH		OTR	15,119	23,930	0	0
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	2,829	292,865	SH		DFND	22	292,865	0	0
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	2,842	294,220	SH		DFND	48	294,220	0	0
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	1	123	SH		OTR	49	12	111	0
AMERICAN AXLE & MFG HOLDINGS	COM	024061103	143	14,803	SH		DFND	49	14,803	0	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	310	7,187	SH		OTR	15,119	7,187	0	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	30,118	697,653	SH		DFND	15	634,134	0	63,519
AMERICAN CAMPUS COMMUNITIES	COM	024835100	7,444	172,436	SH		DFND	22	172,436	0	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	89	2,065	SH		DFND	4	2,065	0	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	27,474	636,418	SH		DFND	48	596,233	0	40,185
AMERICAN CAMPUS COMMUNITIES	COM	024835100	2,879	66,692	SH		DFND	49	66,382	0	310
AMERICAN CAMPUS COMMUNITIES	COM	024835100	5	116	SH		OTR	49	40	76	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	11	248	SH		DFND	59	248	0	0
AMERICAN CAMPUS COMMUNITIES	COM	024835100	0	1	SH		DFND		1	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	24,408	834,751	SH		DFND	15	698,717	0	136,034
AMERICAN EAGLE OUTFITTERS	COM	02553E106	270	9,248	SH		OTR	15,119	9,248	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	4,708	161,002	SH		DFND	22	161,002	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	1,167	39,922	SH		OTR	22,69	39,922	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	112	3,835	SH		DFND	4	3,835	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	19,901	680,592	SH		DFND	48	680,592	0	0
AMERICAN EAGLE OUTFITTERS	COM	02553E106	2,866	98,026	SH		DFND	49	97,726	0	300
AMERICAN EAGLE OUTFITTERS	COM	02553E106	0	1	SH		DFND		1	0	0
AMERICAN ELECTRIC POWER	COM	025537101	1,419	16,749	SH		OTR	15,118	16,749	0	0
AMERICAN ELECTRIC POWER	COM	025537101	2,377	28,058	SH		OTR	15,119	28,058	0	0
AMERICAN ELECTRIC POWER	COM	025537101	3,743	44,187	SH		DFND	15,22	44,187	0	0
AMERICAN ELECTRIC POWER	COM	025537101	179,362	2,117,612	SH		DFND	15	1,282,083	0	835,529
AMERICAN ELECTRIC POWER	COM	025537101	2,351	27,756	SH		OTR	22,80	27,756	0	0
AMERICAN ELECTRIC POWER	COM	025537101	7,028	82,971	SH		DFND	22	82,971	0	0
AMERICAN ELECTRIC POWER	COM	025537101	235	2,772	SH		DFND	4	2,772	0	0
AMERICAN ELECTRIC POWER	COM	025537101	119	1,400	SH		OTR	4	0	1,400	0
AMERICAN ELECTRIC POWER	COM	025537101	114,820	1,355,612	SH		DFND	48	1,206,003	0	149,609
AMERICAN ELECTRIC POWER	COM	025537101	10,761	127,045	SH		DFND	49	126,699	0	346
AMERICAN ELECTRIC POWER	COM	025537101	761	8,981	SH		OTR	49	5,278	3,703	0
AMERICAN ELECTRIC POWER	COM	025537101	2,045	24,146	SH		DFND	59	23,688	0	458
AMERICAN EQUITY INVT LIFE HL	COM	025676206	18,882	598,871	SH		DFND	15	516,482	0	82,389
AMERICAN EQUITY INVT LIFE HL	COM	025676206	267	8,473	SH		OTR	15,119	8,473	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	7,019	222,614	SH		DFND	22	222,614	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	30	950	SH		DFND	4	950	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	7,338	232,742	SH		DFND	48	232,742	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	247	7,826	SH		OTR	49	7,826	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	1,684	53,415	SH		DFND	49	53,415	0	0
AMERICAN EQUITY INVT LIFE HL	COM	025676206	3	89	SH		DFND	59	1	0	88
AMERICAN EXPRESS CO	COM	025816109	3,117	22,036	SH		OTR	15,118	22,036	0	0
AMERICAN EXPRESS CO	COM	025816109	8,335	58,931	SH		DFND	15,22	58,931	0	0
AMERICAN EXPRESS CO	COM	025816109	3,143	22,222	SH		OTR	15,119	22,222	0	0
AMERICAN EXPRESS CO	COM	025816109	419,734	2,967,574	SH		DFND	15	1,863,718	0	1,103,856
AMERICAN EXPRESS CO	COM	025816109	15,677	110,839	SH		DFND	22	110,839	0	0
AMERICAN EXPRESS CO	COM	025816109	3,455	24,430	SH		DFND	4	24,430	0	0
AMERICAN EXPRESS CO	COM	025816109	2,466	17,433	SH		OTR	4	9,818	7,615	0
AMERICAN EXPRESS CO	COM	025816109	47	332	SH		DFND	40	332	0	0
AMERICAN EXPRESS CO	COM	025816109	278,624	1,969,906	SH		DFND	48	1,771,923	0	197,983
AMERICAN EXPRESS CO	COM	025816109	88,484	625,594	SH		DFND	49	603,994	0	21,600
AMERICAN EXPRESS CO	COM	025816109	22,276	157,491	SH		OTR	49	56,492	49,995	51,004
AMERICAN EXPRESS CO	COM	025816109	1,080	7,633	SH		DFND	59	7,109	0	524
AMERICAN FINANCE TRUST INC	COM	02607T109	1,337	136,157	SH		DFND	15	34,405	0	101,752
AMERICAN FINANCE TRUST INC	COM	02607T109	2,696	274,586	SH		DFND	48	274,586	0	0
AMERICAN FINANCIAL GROUP INC	COM	025932104	47,080	412,621	SH		DFND	15	375,884	0	36,737
AMERICAN FINANCIAL GROUP INC	COM	025932104	553	4,850	SH		OTR	15,119	4,850	0	0
AMERICAN FINANCIAL GROUP INC	COM	025932104	9,708	85,083	SH		DFND	22	85,083	0	0
AMERICAN FINANCIAL GROUP INC	COM	025932104	140	1,223	SH		DFND	4	1,223	0	0
AMERICAN FINANCIAL GROUP INC	COM	025932104	30,543	267,686	SH		DFND	48	245,878	0	21,808
AMERICAN FINANCIAL GROUP INC	COM	025932104	8,945	78,396	SH		OTR	49	44	402	77,950
AMERICAN FINANCIAL GROUP INC	COM	025932104	4,913	43,059	SH		DFND	49	42,913	0	146
AMERICAN FINANCIAL GROUP INC	COM	025932104	0	1	SH		DFND	59	1	0	0
AMERICAN FINANCIAL GROUP INC	COM	025932104	0	1	SH		DFND		1	0	0
AMERICAN HOMES 4 RENT- A	CL A	02665T306	12,852	385,490	SH		DFND	15	255,970	0	129,520
AMERICAN HOMES 4 RENT- A	CL A	02665T306	3,040	91,169	SH		OTR	22,80	91,169	0	0
AMERICAN HOMES 4 RENT- A	CL A	02665T306	1,717	51,497	SH		DFND	40	51,497	0	0
AMERICAN HOMES 4 RENT- A	CL A	02665T306	23,956	718,540	SH		DFND	48	636,677	0	81,863
AMERICAN HOMES 4 RENT- A	CL A	02665T306	283	8,502	SH		DFND	49	8,502	0	0
AMERICAN HOMES 4 RENT- A	CL A	02665T306	26	773	SH		OTR	49	607	166	0
AMERICAN HOMES 4 RENT- A	CL A	02665T306	23	701	SH		DFND	59	701	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	3,600	77,905	SH		DFND	15,22	77,905	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	2,858	61,844	SH		OTR	15,119	61,844	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	149,209	3,228,934	SH		DFND	15	1,762,540	0	1,466,394
AMERICAN INTERNATIONAL GROUP	COM	026874784	1,348	29,164	SH		OTR	15,118	29,164	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	3,667	79,361	SH		OTR	22,80	79,361	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	44,087	954,064	SH		DFND	22	954,064	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	442	9,567	SH		OTR	4	4,788	4,680	99
AMERICAN INTERNATIONAL GROUP	COM	026874784	5,495	118,919	SH		DFND	4	118,919	0	0
AMERICAN INTERNATIONAL GROUP	COM	026874784	118,189	2,557,651	SH		DFND	48	2,296,563	0	261,088
AMERICAN INTERNATIONAL GROUP	COM	026874784	183,327	3,967,251	SH		DFND	49	3,925,845	1,450	39,956
AMERICAN INTERNATIONAL GROUP	COM	026874784	7,049	152,536	SH		OTR	49	98,651	53,685	200
AMERICAN INTERNATIONAL GROUP	COM	026874784	6,717	145,364	SH		DFND	59	144,074	0	1,290
AMERICAN NATIONAL GROUP INC	COM	02772A109	9,511	88,168	SH		DFND	15	84,702	0	3,466
AMERICAN NATIONAL GROUP INC	COM	02772A109	1,365	12,655	SH		DFND	48	10,543	0	2,112
AMERICAN NATIONAL GROUP INC	COM	02772A109	257	2,382	SH		DFND	49	2,382	0	0
AMERICAN OUTDOOR BRANDS INC	COM	02875D109	295	11,689	SH		DFND	15	3,184	0	8,505
AMERICAN OUTDOOR BRANDS INC	COM	02875D109	902	35,794	SH		DFND	48	35,794	0	0
AMERICAN OUTDOOR BRANDS INC	COM	02875D109	2	97	SH		DFND	49	97	0	0
AMERICAN PUBLIC EDUCATION	COM	02913V103	3,437	96,453	SH		DFND	15	85,049	0	11,404
AMERICAN PUBLIC EDUCATION	COM	02913V103	1,632	45,797	SH		DFND	22	45,797	0	0
AMERICAN PUBLIC EDUCATION	COM	02913V103	8	225	SH		DFND	4	225	0	0
AMERICAN PUBLIC EDUCATION	COM	02913V103	1,297	36,396	SH		DFND	48	36,396	0	0
AMERICAN PUBLIC EDUCATION	COM	02913V103	59	1,655	SH		DFND	49	1,655	0	0
AMERICAN PUBLIC EDUCATION	COM	02913V103	0	1	SH		DFND		1	0	0
AMERICAN RESOURCES CORP	CL A	02927U208	131	34,588	SH		DFND	48	34,588	0	0
AMERICAN SOFTWARE INC-CL A	CL A	029683109	3,951	190,859	SH		DFND	15	177,232	0	13,627
AMERICAN SOFTWARE INC-CL A	CL A	029683109	1,550	74,897	SH		DFND	48	74,897	0	0
AMERICAN SOFTWARE INC-CL A	CL A	029683109	68	3,286	SH		DFND	49	3,286	0	0
AMERICAN SOFTWARE INC-CL A	CL A	029683109	5	264	SH		DFND	59	264	0	0
AMERICAN STATES WATER CO	COM	029899101	17,187	227,275	SH		DFND	15	213,616	0	13,659
AMERICAN STATES WATER CO	COM	029899101	7,193	95,116	SH		DFND	22	95,116	0	0
AMERICAN STATES WATER CO	COM	029899101	51	675	SH		DFND	4	675	0	0
AMERICAN STATES WATER CO	COM	029899101	7,638	101,004	SH		DFND	48	101,004	0	0
AMERICAN STATES WATER CO	COM	029899101	91	1,197	SH		OTR	49	0	1,197	0
AMERICAN STATES WATER CO	COM	029899101	1,589	21,018	SH		DFND	49	21,018	0	0
AMERICAN SUPERCONDUCTOR CORP	SHS	030111207	807	42,583	SH		DFND	15	16,640	0	25,943
AMERICAN SUPERCONDUCTOR CORP	SHS	030111207	1,557	82,122	SH		DFND	48	82,122	0	0
AMERICAN TOWER CORP	COM	03027X100	9,605	40,178	SH		DFND	15,22	40,178	0	0
AMERICAN TOWER CORP	COM	03027X100	438,973	1,836,245	SH		DFND	15	1,087,049	0	749,196
AMERICAN TOWER CORP	COM	03027X100	1,678	7,019	SH		OTR	15,119	7,019	0	0
AMERICAN TOWER CORP	COM	03027X100	3,588	15,010	SH		OTR	15,118	15,010	0	0
AMERICAN TOWER CORP	COM	03027X100	33,253	139,101	SH		DFND	22	139,101	0	0
AMERICAN TOWER CORP	COM	03027X100	2,801	11,717	SH		DFND	4	11,717	0	0
AMERICAN TOWER CORP	COM	03027X100	4,705	19,680	SH		OTR	4	809	18,034	837
AMERICAN TOWER CORP	COM	03027X100	26	108	SH		DFND	40	108	0	0
AMERICAN TOWER CORP	COM	03027X100	318,176	1,330,946	SH		DFND	48	1,194,060	0	136,886
AMERICAN TOWER CORP	COM	03027X100	11,807	49,391	SH		OTR	49	14,799	31,741	2,851
AMERICAN TOWER CORP	COM	03027X100	125,602	525,400	SH		DFND	49	519,696	74	5,630
AMERICAN TOWER CORP	COM	03027X100	18,319	76,630	SH		DFND	59	73,283	0	3,347
AMERICAN VANGUARD CORP	COM	030371108	2,746	134,533	SH		DFND	15	106,619	0	27,914
AMERICAN VANGUARD CORP	COM	030371108	2,514	123,191	SH		OTR	22,121	123,191	0	0
AMERICAN VANGUARD CORP	COM	030371108	1,369	67,095	SH		DFND	22	67,095	0	0
AMERICAN VANGUARD CORP	COM	030371108	1,425	69,840	SH		DFND	48	69,840	0	0
AMERICAN VANGUARD CORP	COM	030371108	57	2,770	SH		DFND	49	2,770	0	0
AMERICAN VANGUARD CORP	COM	030371108	0	1	SH		DFND		1	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	916	6,111	SH		OTR	15,118	6,111	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	2,476	16,513	SH		DFND	15,22	16,513	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	640	4,271	SH		OTR	15,119	4,271	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	96,232	641,886	SH		DFND	15	349,962	0	291,924
AMERICAN WATER WORKS CO INC	COM	030420103	8,766	58,473	SH		DFND	22	58,473	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	159	1,063	SH		DFND	4	1,063	0	0
AMERICAN WATER WORKS CO INC	COM	030420103	106,649	711,376	SH		DFND	48	656,791	0	54,585
AMERICAN WATER WORKS CO INC	COM	030420103	5,560	37,087	SH		DFND	49	37,063	0	24
AMERICAN WATER WORKS CO INC	COM	030420103	314	2,096	SH		OTR	49	287	1,809	0
AMERICAN WATER WORKS CO INC	COM	030420103	0	1	SH		DFND		1	0	0
AMERICAN WELL CORP-CLASS A	CL A	03044L105	836	48,111	SH		DFND	15	26,491	0	21,620
AMERICAN WELL CORP-CLASS A	CL A	03044L105	939	54,081	SH		DFND	15,22	54,081	0	0
AMERICAN WELL CORP-CLASS A	CL A	03044L105	63	3,601	SH		DFND	22	3,601	0	0
AMERICAN WELL CORP-CLASS A	CL A	03044L105	1,675	96,441	SH		DFND	48	80,103	0	16,338
AMERICAN WELL CORP-CLASS A	CL A	03044L105	21	1,230	SH		DFND	49	1,230	0	0
AMERICAN WOODMARK CORP	COM	030506109	8,715	88,405	SH		DFND	15	72,559	0	15,846
AMERICAN WOODMARK CORP	COM	030506109	4,278	43,400	SH		DFND	22	43,400	0	0
AMERICAN WOODMARK CORP	COM	030506109	2	25	SH		DFND	4	25	0	0
AMERICAN WOODMARK CORP	COM	030506109	4,231	42,915	SH		DFND	48	42,915	0	0
AMERICAN WOODMARK CORP	COM	030506109	440	4,464	SH		DFND	49	4,464	0	0
AMERICAN WOODMARK CORP	COM	030506109	61	620	SH		OTR	49	0	620	0
AMERICAN WOODMARK CORP	COM	030506109	0	1	SH		DFND		1	0	0
AMERICA'S CAR-MART INC	COM	03062T105	5,139	33,726	SH		DFND	15	31,017	0	2,709
AMERICA'S CAR-MART INC	COM	03062T105	2,367	15,533	SH		DFND	22	15,533	0	0
AMERICA'S CAR-MART INC	COM	03062T105	9	60	SH		DFND	4	60	0	0
AMERICA'S CAR-MART INC	COM	03062T105	2,620	17,196	SH		DFND	48	17,196	0	0
AMERICA'S CAR-MART INC	COM	03062T105	130	850	SH		DFND	49	850	0	0
AMERICA'S CAR-MART INC	COM	03062T105	228	1,494	SH		DFND	59	1,494	0	0
AMERICOLD REALTY TRUST	COM	03064D108	34,797	904,519	SH		DFND	15,22	904,519	0	0
AMERICOLD REALTY TRUST	COM	03064D108	22,546	586,077	SH		DFND	15	454,593	0	131,484
AMERICOLD REALTY TRUST	COM	03064D108	5,806	150,926	SH		DFND	22	150,926	0	0
AMERICOLD REALTY TRUST	COM	03064D108	23	590	SH		DFND	4	590	0	0
AMERICOLD REALTY TRUST	COM	03064D108	845	21,975	SH		DFND	42	21,975	0	0
AMERICOLD REALTY TRUST	COM	03064D108	25,612	665,765	SH		DFND	48	595,299	0	70,466
AMERICOLD REALTY TRUST	COM	03064D108	1,517	39,446	SH		DFND	49	39,446	0	0
AMERICOLD REALTY TRUST	COM	03064D108	355	9,232	SH		OTR	49	7,520	1,712	0
AMERICOLD REALTY TRUST	COM	03064D108	89	2,313	SH		DFND	59	2,313	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	11,302	48,623	SH		OTR	15,118	48,623	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	2,460	10,581	SH		DFND	15,22	10,581	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	1,060	4,561	SH		OTR	15,119	4,561	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	274,084	1,179,110	SH		DFND	15	954,932	0	224,178
AMERIPRISE FINANCIAL INC	COM	03076C106	11,000	47,321	SH		OTR	22,80	47,321	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	67,809	291,715	SH		DFND	22	291,715	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	601	2,584	SH		OTR	4	816	1,768	0
AMERIPRISE FINANCIAL INC	COM	03076C106	2,907	12,506	SH		DFND	4	12,506	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	259	1,113	SH		DFND	42	1,113	0	0
AMERIPRISE FINANCIAL INC	COM	03076C106	91,598	394,053	SH		DFND	48	358,428	0	35,625
AMERIPRISE FINANCIAL INC	COM	03076C106	10,977	47,225	SH		OTR	49	19,093	26,598	1,534
AMERIPRISE FINANCIAL INC	COM	03076C106	109,638	471,664	SH		DFND	49	464,910	67	6,687
AMERIPRISE FINANCIAL INC	COM	03076C106	729	3,136	SH		DFND	59	3,116	0	20
AMERIPRISE FINANCIAL INC	COM	03076C106	0	1	SH		DFND		1	0	0
AMERIS BANCORP	COM	03076K108	21,706	413,372	SH		DFND	15	354,598	0	58,774
AMERIS BANCORP	COM	03076K108	9,464	180,227	SH		DFND	22	180,227	0	0
AMERIS BANCORP	COM	03076K108	188	3,575	SH		DFND	4	3,575	0	0
AMERIS BANCORP	COM	03076K108	8,739	166,425	SH		DFND	48	166,425	0	0
AMERIS BANCORP	COM	03076K108	3,349	63,779	SH		DFND	49	63,779	0	0
AMERISAFE INC	COM	03071H100	9,551	149,237	SH		DFND	15	131,717	0	17,520
AMERISAFE INC	COM	03071H100	3,259	50,918	SH		DFND	22	50,918	0	0
AMERISAFE INC	COM	03071H100	7	108	SH		DFND	4	108	0	0
AMERISAFE INC	COM	03071H100	3,019	47,174	SH		DFND	48	47,174	0	0
AMERISAFE INC	COM	03071H100	698	10,910	SH		DFND	49	10,910	0	0
AMERISAFE INC	COM	03071H100	159	2,478	SH		DFND	59	2,361	0	117
AMERISERV FINANCIAL INC	COM	03074A102	43	10,604	SH		DFND	48	10,604	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	1,686	14,277	SH		OTR	15,119	14,277	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	1,550	13,128	SH		DFND	15,22	13,128	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	600	5,081	SH		OTR	15,118	5,081	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	98,641	835,446	SH		DFND	15	594,773	0	240,673
AMERISOURCEBERGEN CORP	COM	03073E105	4,733	40,090	SH		OTR	22,80	40,090	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	7,971	67,512	SH		DFND	22	67,512	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	128	1,086	SH		DFND	4	1,086	0	0
AMERISOURCEBERGEN CORP	COM	03073E105	6	47	SH		OTR	4	0	0	47
AMERISOURCEBERGEN CORP	COM	03073E105	53,114	449,851	SH		DFND	48	405,426	0	44,425
AMERISOURCEBERGEN CORP	COM	03073E105	6,238	52,837	SH		DFND	49	52,731	0	106
AMERISOURCEBERGEN CORP	COM	03073E105	477	4,043	SH		OTR	49	1,597	2,446	0
AMERISOURCEBERGEN CORP	COM	03073E105	4,884	41,362	SH		DFND	59	39,968	0	1,395
AMERISOURCEBERGEN CORP	COM	03073E105	0	1	SH		DFND		1	0	0
AMES NATIONAL CORP	COM	031001100	916	35,804	SH		DFND	15	27,797	0	8,007
AMES NATIONAL CORP	COM	031001100	570	22,291	SH		DFND	48	22,291	0	0
AMES NATIONAL CORP	COM	031001100	13	500	SH		DFND	49	500	0	0
AMETEK INC	COM	031100100	992	7,768	SH		OTR	15,118	7,768	0	0
AMETEK INC	COM	031100100	513	4,018	SH		OTR	15,119	4,018	0	0
AMETEK INC	COM	031100100	158,960	1,244,498	SH		DFND	15	875,542	0	368,956
AMETEK INC	COM	031100100	2,667	20,878	SH		DFND	15,22	20,878	0	0
AMETEK INC	COM	031100100	7,663	59,994	SH		OTR	22,80	59,994	0	0
AMETEK INC	COM	031100100	73,566	575,952	SH		DFND	22	575,952	0	0
AMETEK INC	COM	031100100	181	1,419	SH		DFND	4	1,419	0	0
AMETEK INC	COM	031100100	1,250	9,789	SH		OTR	4	31	9,710	48
AMETEK INC	COM	031100100	115	898	SH		DFND	42	898	0	0
AMETEK INC	COM	031100100	88,732	694,688	SH		DFND	48	625,591	0	69,097
AMETEK INC	COM	031100100	9,210	72,108	SH		DFND	49	71,491	0	617
AMETEK INC	COM	031100100	355	2,782	SH		OTR	49	1,582	1,170	30
AMETEK INC	COM	031100100	937	7,337	SH		DFND	59	7,171	0	166
AMGEN INC	COM	031162100	12,900	51,847	SH		DFND	15,22	51,847	0	0
AMGEN INC	COM	031162100	4,526	18,189	SH		OTR	15,119	18,189	0	0
AMGEN INC	COM	031162100	4,855	19,512	SH		OTR	15,118	19,512	0	0
AMGEN INC	COM	031162100	957,932	3,850,053	SH		DFND	15	2,869,262	0	980,791
AMGEN INC	COM	031162100	27,572	110,816	SH		DFND	22	110,816	0	0
AMGEN INC	COM	031162100	1,585	6,371	SH		DFND	4	6,371	0	0
AMGEN INC	COM	031162100	1,128	4,534	SH		OTR	4	387	4,138	9
AMGEN INC	COM	031162100	59	238	SH		DFND	40	238	0	0
AMGEN INC	COM	031162100	439,595	1,766,791	SH		DFND	48	1,585,999	0	180,792
AMGEN INC	COM	031162100	114,343	459,559	SH		DFND	49	448,175	0	11,384
AMGEN INC	COM	031162100	21,236	85,351	SH		OTR	49	41,621	43,029	701
AMGEN INC	COM	031162100	5,464	21,960	SH		DFND	59	21,184	0	776
AMGEN INC	COM	031162100	122	491	SH		DFND		491	0	0
AMICUS THERAPEUTICS INC	COM	03152W109	3,261	330,099	SH		DFND	15	237,874	0	92,225
AMICUS THERAPEUTICS INC	COM	03152W109	516	52,232	SH		OTR	22,87	52,232	0	0
AMICUS THERAPEUTICS INC	COM	03152W109	6,481	656,011	SH		DFND	48	656,011	0	0
AMICUS THERAPEUTICS INC	COM	03152W109	87	8,822	SH		DFND	49	8,822	0	0
AMKOR TECHNOLOGY INC	COM	031652100	19,287	813,453	SH		DFND	15	735,281	0	78,172
AMKOR TECHNOLOGY INC	COM	031652100	20	856	SH		OTR	15,119	856	0	0
AMKOR TECHNOLOGY INC	COM	031652100	2,730	115,159	SH		DFND	22	115,159	0	0
AMKOR TECHNOLOGY INC	COM	031652100	12,145	512,247	SH		DFND	48	512,247	0	0
AMKOR TECHNOLOGY INC	COM	031652100	821	34,612	SH		OTR	49	0	34,612	0
AMKOR TECHNOLOGY INC	COM	031652100	19	820	SH		DFND	49	820	0	0
AMKOR TECHNOLOGY INC	COM	031652100	0	1	SH		DFND		1	0	0
AMMO INC	COM	00175J107	120	20,298	SH		DFND	48	20,298	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	17,776	241,191	SH		DFND	15	223,956	0	17,235
AMN HEALTHCARE SERVICES INC	COM	001744101	157	2,135	SH		OTR	22,87	2,135	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	8,980	121,843	SH		DFND	22	121,843	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	2,202	29,879	SH		OTR	22,96	29,879	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	2,324	31,532	SH		OTR	22,114	31,532	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	59	805	SH		DFND	4	805	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	9,285	125,988	SH		DFND	48	125,988	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	1,401	19,010	SH		DFND	49	19,010	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	503	6,831	SH		OTR	49	6,831	0	0
AMN HEALTHCARE SERVICES INC	COM	001744101	627	8,507	SH		DFND	59	8,171	0	336
AMN HEALTHCARE SERVICES INC	COM	001744101	0	1	SH		DFND		1	0	0
AMNEAL PHARMACEUTICALS INC	COM	03168L105	2,491	370,102	SH		DFND	15	338,286	0	31,816
AMNEAL PHARMACEUTICALS INC	COM	03168L105	1,813	269,441	SH		DFND	48	269,441	0	0
AMPCO-PITTSBURGH CORP	COM	032037103	49	7,200	SH		DFND	15	7,200	0	0
AMPCO-PITTSBURGH CORP	COM	032037103	125	18,490	SH		OTR	22,83	18,490	0	0
AMPCO-PITTSBURGH CORP	COM	032037103	136	20,170	SH		DFND	48	20,170	0	0
AMPHASTAR PHARMACEUTICALS IN	COM	03209R103	3,322	181,351	SH		DFND	15	168,977	0	12,374
AMPHASTAR PHARMACEUTICALS IN	COM	03209R103	1,786	97,505	SH		DFND	22	97,505	0	0
AMPHASTAR PHARMACEUTICALS IN	COM	03209R103	12	656	SH		DFND	4	656	0	0
AMPHASTAR PHARMACEUTICALS IN	COM	03209R103	1,650	90,049	SH		DFND	48	90,049	0	0
AMPHASTAR PHARMACEUTICALS IN	COM	03209R103	167	9,122	SH		DFND	49	9,122	0	0
AMPHENOL CORP-CL A	CL A	032095101	674	10,216	SH		OTR	15,119	10,216	0	0
AMPHENOL CORP-CL A	CL A	032095101	3,549	53,791	SH		DFND	15,22	53,791	0	0
AMPHENOL CORP-CL A	CL A	032095101	176,337	2,672,994	SH		DFND	15	1,686,119	0	986,875
AMPHENOL CORP-CL A	CL A	032095101	1,333	20,204	SH		OTR	15,118	20,204	0	0
AMPHENOL CORP-CL A	CL A	032095101	19,401	294,082	SH		DFND	22	294,082	0	0
AMPHENOL CORP-CL A	CL A	032095101	11,431	173,270	SH		OTR	22,71	173,270	0	0
AMPHENOL CORP-CL A	CL A	032095101	449	6,808	SH		DFND	4	6,808	0	0
AMPHENOL CORP-CL A	CL A	032095101	91	1,380	SH		OTR	4	1,060	320	0
AMPHENOL CORP-CL A	CL A	032095101	117,386	1,779,378	SH		DFND	48	1,600,071	0	179,307
AMPHENOL CORP-CL A	CL A	032095101	20,388	309,044	SH		DFND	49	306,951	0	2,093
AMPHENOL CORP-CL A	CL A	032095101	1,707	25,868	SH		OTR	49	11,128	14,740	0
AMPHENOL CORP-CL A	CL A	032095101	1,342,615	20,351,908	SH		DFND	51	14,113,684	0	6,238,224
AMPHENOL CORP-CL A	CL A	032095101	56,774	860,600	SH		DFND	51,22	860,600	0	0
AMPHENOL CORP-CL A	CL A	032095101	175,414	2,659,004	SH		OTR	51,94	2,659,004	0	0
AMPHENOL CORP-CL A	CL A	032095101	3,747	56,800	SH		OTR	51	56,800	0	0
AMPHENOL CORP-CL A	CL A	032095101	26,180	396,844	SH		OTR	51,48	396,844	0	0
AMPHENOL CORP-CL A	CL A	032095101	549	8,322	SH		DFND	59	8,280	0	42
AMPHENOL CORP-CL A	CL A	032095101	396	6,000	SH		DFND		6,000	0	0
AMPIO PHARMACEUTICALS INC	COM	03209T109	179	105,668	SH		DFND	48	105,668	0	0
AMPLIFY ENERGY CORP	COM	03212B103	113	40,824	SH		DFND	48	40,824	0	0
AMPLIFY ENERGY CORP	COM	03212B103	0	4	SH		DFND		4	0	0
AMTECH SYSTEMS INC	COM	032332504	206	17,442	SH		DFND	48	17,442	0	0
AMYRIS INC	COM	03236M200	2,360	123,536	SH		DFND	15	37,931	0	85,605
AMYRIS INC	COM	03236M200	1,157	60,585	SH		OTR	22,69	60,585	0	0
AMYRIS INC	COM	03236M200	656	34,326	SH		OTR	22,78	34,326	0	0
AMYRIS INC	COM	03236M200	5,556	290,907	SH		DFND	48	290,907	0	0
AMYRIS INC	COM	03236M200	79	4,130	SH		OTR	49	0	4,130	0
ANALOG DEVICES INC	COM	032654105	5,115	32,981	SH		DFND	15,22	32,981	0	0
ANALOG DEVICES INC	COM	032654105	272,042	1,754,206	SH		DFND	15	1,131,337	0	622,869
ANALOG DEVICES INC	COM	032654105	1,931	12,449	SH		OTR	15,118	12,449	0	0
ANALOG DEVICES INC	COM	032654105	1,095	7,060	SH		OTR	15,119	7,060	0	0
ANALOG DEVICES INC	COM	032654105	9,634	62,120	SH		DFND	22	62,120	0	0
ANALOG DEVICES INC	COM	032654105	2,039	13,151	SH		OTR	4	35	13,100	16
ANALOG DEVICES INC	COM	032654105	594	3,830	SH		DFND	4	3,830	0	0
ANALOG DEVICES INC	COM	032654105	167,230	1,078,344	SH		DFND	48	966,879	0	111,465
ANALOG DEVICES INC	COM	032654105	3,404	21,952	SH		OTR	49	8,426	13,301	225
ANALOG DEVICES INC	COM	032654105	36,005	232,167	SH		DFND	49	229,296	0	2,871
ANALOG DEVICES INC	COM	032654105	845	5,448	SH		DFND	59	5,301	0	147
ANALOG DEVICES INC	COM	032654105	0	1	SH		DFND		1	0	0
ANAPLAN INC	COM	03272L108	24,481	454,613	SH		DFND	15	392,254	0	62,359
ANAPLAN INC	COM	03272L108	392	7,275	SH		DFND	22	7,275	0	0
ANAPLAN INC	COM	03272L108	4	65	SH		DFND	4	65	0	0
ANAPLAN INC	COM	03272L108	16,449	305,462	SH		DFND	48	264,722	0	40,740
ANAPLAN INC	COM	03272L108	2	38	SH		OTR	49	0	38	0
ANAPLAN INC	COM	03272L108	115	2,140	SH		DFND	49	2,140	0	0
ANAPLAN INC	COM	03272L108	38	705	SH		DFND	59	554	0	151
ANAPTYSBIO INC	COM	032724106	936	43,453	SH		DFND	15	24,608	0	18,845
ANAPTYSBIO INC	COM	032724106	1,179	54,691	SH		DFND	48	54,691	0	0
ANAPTYSBIO INC	COM	032724106	49	2,285	SH		DFND	49	2,285	0	0
ANAVEX LIFE SCIENCES CORP	COM	032797300	506	33,851	SH		DFND	15	7,402	0	26,449
ANAVEX LIFE SCIENCES CORP	COM	032797300	2,227	148,975	SH		DFND	48	148,975	0	0
ANAVEX LIFE SCIENCES CORP	COM	032797300	78	5,213	SH		DFND	49	5,213	0	0
ANAVEX LIFE SCIENCES CORP	COM	032797300	251	16,783	SH		DFND	59	16,783	0	0
ANAVEX LIFE SCIENCES CORP	COM	032797300	0	1	SH		DFND		1	0	0
ANDERSONS INC/THE	COM	034164103	8,008	292,484	SH		DFND	15	264,257	0	28,227
ANDERSONS INC/THE	COM	034164103	2,158	78,831	SH		DFND	22	78,831	0	0
ANDERSONS INC/THE	COM	034164103	1,045	38,150	SH		OTR	22,83	38,150	0	0
ANDERSONS INC/THE	COM	034164103	2,100	76,692	SH		DFND	48	76,692	0	0
ANDERSONS INC/THE	COM	034164103	273	9,970	SH		DFND	49	9,970	0	0
ANDERSONS INC/THE	COM	034164103	0	1	SH		DFND		1	0	0
ANGI INC	COM	00183L102	550	42,312	SH		DFND	15	42,312	0	0
ANGI INC	COM	00183L102	683	52,536	SH		DFND	48	52,536	0	0
ANGI INC	COM	00183L102	28	2,184	SH		OTR	49	0	2,184	0
ANGI INC	COM	00183L102	1	45	SH		DFND	49	45	0	0
ANGIODYNAMICS INC	COM	03475V101	4,506	192,585	SH		DFND	15	156,608	0	35,977
ANGIODYNAMICS INC	COM	03475V101	2,275	97,217	SH		DFND	22	97,217	0	0
ANGIODYNAMICS INC	COM	03475V101	1,322	56,481	SH		OTR	22,121	56,481	0	0
ANGIODYNAMICS INC	COM	03475V101	2,212	94,530	SH		DFND	48	94,530	0	0
ANGIODYNAMICS INC	COM	03475V101	123	5,250	SH		DFND	49	5,250	0	0
ANGIODYNAMICS INC	COM	03475V101	0	1	SH		DFND		1	0	0
ANGION BIOMEDICA CORP	COM	03476J107	19	1,078	SH		DFND	15	0	0	1,078
ANGION BIOMEDICA CORP	COM	03476J107	169	9,339	SH		DFND	48	9,339	0	0
ANHEUSER-BUSCH INBEV-SPN ADR	SPON	03524A108	20	320	SH		DFND	4	320	0	0
ANHEUSER-BUSCH INBEV-SPN ADR	SPON	03524A108	17	274	SH		OTR	49	126	148	0
ANHEUSER-BUSCH INBEV-SPN ADR	SPON	03524A108	823	13,096	SH		DFND	49	13,096	0	0
ANHEUSER-BUSCH INBEV-SPN ADR	SPON	03524A108	1,245	19,815	SH		DFND	59	19,105	0	710
ANHEUSER-BUSCH INBEV-SPN ADR	SPON	03524A108	0	1	SH		DFND		1	0	0
ANI PHARMACEUTICALS INC	COM	00182C103	1,667	46,113	SH		DFND	15	39,308	0	6,805
ANI PHARMACEUTICALS INC	COM	00182C103	951	26,324	SH		DFND	22	26,324	0	0
ANI PHARMACEUTICALS INC	COM	00182C103	936	25,897	SH		DFND	48	25,897	0	0
ANI PHARMACEUTICALS INC	COM	00182C103	92	2,545	SH		DFND	49	2,545	0	0
ANIKA THERAPEUTICS INC	COM	035255108	3,248	79,619	SH		DFND	15	66,137	0	13,482
ANIKA THERAPEUTICS INC	COM	035255108	1,531	37,537	SH		DFND	22	37,537	0	0
ANIKA THERAPEUTICS INC	COM	035255108	1,595	39,093	SH		DFND	48	39,093	0	0
ANIKA THERAPEUTICS INC	COM	035255108	64	1,560	SH		DFND	49	1,560	0	0
ANIXA BIOSCIENCES INC	COM	03528H109	229	48,942	SH		DFND	48	48,942	0	0
ANIXA BIOSCIENCES INC	COM	03528H109	9	2,000	SH		DFND	49	2,000	0	0
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	741	86,171	SH		OTR	15,119	86,171	0	0
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	18,250	2,122,053	SH		DFND	15	1,390,743	0	731,310
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	872	101,340	SH		DFND	22	101,340	0	0
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	26	3,000	SH		OTR	4	0	0	3,000
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	18,782	2,183,974	SH		DFND	48	1,754,268	0	429,706
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	661	76,862	SH		OTR	49	75,620	1,242	0
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	468	54,370	SH		DFND	49	54,370	0	0
ANNALY CAPITAL MANAGEMENT IN	COM	035710409	316	36,802	SH		DFND	59	36,173	0	629
ANNEXON INC	COM	03589W102	641	23,039	SH		DFND	15	12,978	0	10,061
ANNEXON INC	COM	03589W102	1,543	55,431	SH		DFND	48	55,431	0	0
ANSYS INC	COM	03662Q105	281	827	SH		OTR	15,119	827	0	0
ANSYS INC	COM	03662Q105	2,634	7,756	SH		DFND	15,22	7,756	0	0
ANSYS INC	COM	03662Q105	1,008	2,968	SH		OTR	15,118	2,968	0	0
ANSYS INC	COM	03662Q105	249,224	733,963	SH		DFND	15	567,674	0	166,289
ANSYS INC	COM	03662Q105	52,531	154,702	SH		DFND	22	154,702	0	0
ANSYS INC	COM	03662Q105	12,512	36,848	SH		OTR	22,71	36,848	0	0
ANSYS INC	COM	03662Q105	3	9	SH		OTR	4	9	0	0
ANSYS INC	COM	03662Q105	370	1,090	SH		DFND	4	1,090	0	0
ANSYS INC	COM	03662Q105	72	212	SH		DFND	42	212	0	0
ANSYS INC	COM	03662Q105	89,323	263,054	SH		DFND	48	236,545	0	26,509
ANSYS INC	COM	03662Q105	1,151	3,389	SH		OTR	49	547	2,842	0
ANSYS INC	COM	03662Q105	13,919	40,992	SH		DFND	49	40,796	0	196
ANSYS INC	COM	03662Q105	8,228	24,230	SH		DFND	51	24,230	0	0
ANSYS INC	COM	03662Q105	12,801	37,700	SH		DFND	51,22	37,700	0	0
ANSYS INC	COM	03662Q105	2,195	6,463	SH		DFND	59	6,251	0	212
ANSYS INC	COM	03662Q105	0	1	SH		DFND		1	0	0
ANTARES PHARMA INC	COM	036642106	1,078	262,233	SH		DFND	15	214,058	0	48,175
ANTARES PHARMA INC	COM	036642106	1,703	414,472	SH		DFND	48	414,472	0	0
ANTERIX INC	COM	03676C100	357	7,565	SH		DFND	15	3,641	0	3,924
ANTERIX INC	COM	03676C100	1,377	29,190	SH		DFND	48	29,190	0	0
ANTERO MIDSTREAM CORP	COM	03676B102	109,216	12,094,824	SH		DFND	15	11,973,639	0	121,185
ANTERO MIDSTREAM CORP	COM	03676B102	20	2,247	SH		OTR	15,119	2,247	0	0
ANTERO MIDSTREAM CORP	COM	03676B102	158	17,510	SH		DFND	15,22	17,510	0	0
ANTERO MIDSTREAM CORP	COM	03676B102	3,016	333,962	SH		DFND	22	333,962	0	0
ANTERO MIDSTREAM CORP	COM	03676B102	10,071	1,115,331	SH		DFND	48	1,034,872	0	80,459
ANTERO MIDSTREAM CORP	COM	03676B102	57	6,290	SH		DFND	49	6,290	0	0
ANTERO MIDSTREAM CORP	COM	03676B102	353	39,049	SH		DFND	59	38,570	0	479
ANTERO MIDSTREAM CORP	COM	03676B102	0	1	SH		DFND		1	0	0
ANTERO RESOURCES CORP	COM	03674X106	6,382	625,724	SH		DFND	15	405,985	0	219,739
ANTERO RESOURCES CORP	COM	03674X106	467	45,765	SH		OTR	15,119	45,765	0	0
ANTERO RESOURCES CORP	COM	03674X106	6,754	662,183	SH		DFND	48	662,183	0	0
ANTERO RESOURCES CORP	COM	03674X106	1	144	SH		OTR	49	36	108	0
ANTERO RESOURCES CORP	COM	03674X106	66	6,493	SH		DFND	49	6,249	0	244
ANTHEM INC	COM	036752103	4,713	13,129	SH		OTR	15,119	13,129	0	0
ANTHEM INC	COM	036752103	7,068	19,690	SH		OTR	15,118	19,690	0	0
ANTHEM INC	COM	036752103	7,892	21,986	SH		DFND	15,22	21,986	0	0
ANTHEM INC	COM	036752103	418,107	1,164,807	SH		DFND	15	746,761	0	418,046
ANTHEM INC	COM	036752103	15,249	42,481	SH		DFND	22	42,481	0	0
ANTHEM INC	COM	036752103	536	1,492	SH		DFND	4	1,492	0	0
ANTHEM INC	COM	036752103	58	162	SH		OTR	4	8	129	25
ANTHEM INC	COM	036752103	268,221	747,239	SH		DFND	48	671,409	0	75,830
ANTHEM INC	COM	036752103	28,922	80,573	SH		DFND	49	79,828	0	745
ANTHEM INC	COM	036752103	2,960	8,245	SH		OTR	49	1,035	7,204	6
ANTHEM INC	COM	036752103	1,143	3,185	SH		DFND	59	2,891	0	294
AON PLC-CLASS A	SHS	G0403H108	179,574	780,384	SH		DFND	15	397,681	0	382,703
AON PLC-CLASS A	SHS	G0403H108	1,755	7,625	SH		OTR	15,118	7,625	0	0
AON PLC-CLASS A	SHS	G0403H108	4,690	20,381	SH		DFND	15,22	20,381	0	0
AON PLC-CLASS A	SHS	G0403H108	883	3,837	SH		OTR	15,119	3,837	0	0
AON PLC-CLASS A	SHS	G0403H108	8,932	38,817	SH		DFND	22	38,817	0	0
AON PLC-CLASS A	SHS	G0403H108	3,552	15,434	SH		OTR	22,80	15,434	0	0
AON PLC-CLASS A	SHS	G0403H108	1,021	4,435	SH		DFND	4	4,435	0	0
AON PLC-CLASS A	SHS	G0403H108	59	257	SH		OTR	4	70	0	187
AON PLC-CLASS A	SHS	G0403H108	156,490	680,068	SH		DFND	48	609,391	0	70,677
AON PLC-CLASS A	SHS	G0403H108	3,187	13,849	SH		OTR	49	5,683	8,166	0
AON PLC-CLASS A	SHS	G0403H108	28,663	124,560	SH		DFND	49	119,654	0	4,906
AON PLC-CLASS A	SHS	G0403H108	6,926	30,098	SH		DFND	59	29,258	0	840
APA CORP	COM	03743Q108	236	13,199	SH		OTR	15,118	13,199	0	0
APA CORP	COM	03743Q108	606	33,862	SH		DFND	15,22	33,862	0	0
APA CORP	COM	03743Q108	405	22,653	SH		OTR	15,119	22,653	0	0
APA CORP	COM	03743Q108	30,869	1,724,518	SH		DFND	15	1,127,732	0	596,786
APA CORP	COM	03743Q108	1,119	62,493	SH		DFND	22	62,493	0	0
APA CORP	COM	03743Q108	16	920	SH		DFND	4	920	0	0
APA CORP	COM	03743Q108	21,374	1,194,093	SH		DFND	48	1,082,768	0	111,325
APA CORP	COM	03743Q108	25	1,372	SH		OTR	49	60	1,124	188
APA CORP	COM	03743Q108	1,211	67,671	SH		DFND	49	67,671	0	0
APA CORP	COM	03743Q108	0	1	SH		DFND	59	1	0	0
APA CORP	COM	03743Q108	0	1	SH		DFND		1	0	0
APARTMENT INCOME REIT CO	COM	03750L109	212	4,952	SH		OTR	15,119	4,952	0	0
APARTMENT INCOME REIT CO	COM	03750L109	22,694	530,720	SH		DFND	15	457,843	0	72,877
APARTMENT INCOME REIT CO	COM	03750L109	7,593	177,575	SH		DFND	22	177,575	0	0
APARTMENT INCOME REIT CO	COM	03750L109	78	1,835	SH		DFND	4	1,835	0	0
APARTMENT INCOME REIT CO	COM	03750L109	29,100	680,551	SH		DFND	48	636,019	0	44,532
APARTMENT INCOME REIT CO	COM	03750L109	1,672	39,107	SH		DFND	49	38,827	0	280
APARTMENT INCOME REIT CO	COM	03750L109	19	439	SH		OTR	49	0	439	0
APARTMENT INVT & MGMT CO -A	CL A	03748R747	1,044	169,975	SH		DFND	15	98,139	0	71,836
APARTMENT INVT & MGMT CO -A	CL A	03748R747	10	1,679	SH		DFND	4	1,679	0	0
APARTMENT INVT & MGMT CO -A	CL A	03748R747	2,280	371,334	SH		DFND	48	326,802	0	44,532
APARTMENT INVT & MGMT CO -A	CL A	03748R747	3	476	SH		OTR	49	38	438	0
APARTMENT INVT & MGMT CO -A	CL A	03748R747	53	8,672	SH		DFND	49	8,672	0	0
APELLIS PHARMACEUTICALS INC	COM	03753U106	3,412	79,511	SH		DFND	15	57,137	0	22,374
APELLIS PHARMACEUTICALS INC	COM	03753U106	6,321	147,315	SH		DFND	48	147,315	0	0
APELLIS PHARMACEUTICALS INC	COM	03753U106	13	308	SH		DFND	49	308	0	0
APHRIA INC	COM	03765K104	163	8,863	SH		DFND	48	8,863	0	0
APHRIA INC	COM	03765K104	83	4,500	SH		DFND	49	4,500	0	0
APHRIA INC	COM	03765K104	0	1	SH		DFND		1	0	0
API GROUP CORP	COM	00187Y100	3,532	170,813	SH		DFND	15	43,502	0	127,311
API GROUP CORP	COM	00187Y100	6,812	329,405	SH		DFND	48	329,405	0	0
API GROUP CORP	COM	00187Y100	20	950	SH		DFND	49	950	0	0
API GROUP CORP	COM	00187Y100	72	3,501	SH		DFND	59	3,501	0	0
APOGEE ENTERPRISES INC	COM	037598109	11,388	278,569	SH		DFND	15	258,564	0	20,005
APOGEE ENTERPRISES INC	COM	037598109	2,749	67,241	SH		DFND	22	67,241	0	0
APOGEE ENTERPRISES INC	COM	037598109	8	195	SH		DFND	4	195	0	0
APOGEE ENTERPRISES INC	COM	037598109	2,682	65,601	SH		DFND	48	65,601	0	0
APOGEE ENTERPRISES INC	COM	037598109	1,729	42,305	SH		DFND	49	42,305	0	0
APOGEE ENTERPRISES INC	COM	037598109	0	1	SH		DFND		1	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	137	9,776	SH		OTR	15,119	9,776	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	9,392	672,312	SH		DFND	15	542,062	0	130,250
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	4,682	335,138	SH		DFND	22	335,138	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	4,798	343,480	SH		DFND	48	343,480	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	255	18,263	SH		DFND	49	18,263	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	854	61,118	SH		DFND	59	61,118	0	0
APOLLO COMMERCIAL REAL ESTAT	COM	03762U105	0	1	SH		DFND		1	0	0
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	11,095	236,020	SH		DFND	15	149,016	0	87,004
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	123	2,624	SH		OTR	15,119	2,624	0	0
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	1,074	22,850	SH		DFND	22	22,850	0	0
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	13,588	289,053	SH		DFND	48	237,683	0	51,370
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	209	4,455	SH		DFND	49	4,455	0	0
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	500	10,643	SH		OTR	49	8,471	2,172	0
APOLLO GLOBAL MANAGEMENT INC	COM	03768E105	5,634	119,844	SH		DFND	59	118,520	0	1,324
APOLLO MEDICAL HOLDINGS INC	COM	03763A207	1,722	63,557	SH		DFND	15	56,649	0	6,908
APOLLO MEDICAL HOLDINGS INC	COM	03763A207	1,229	45,350	SH		DFND	48	45,350	0	0
APPFOLIO INC - A	COM	03783C100	22,560	159,533	SH		DFND	15	154,011	0	5,522
APPFOLIO INC - A	COM	03783C100	5,819	41,147	SH		DFND	48	41,147	0	0
APPFOLIO INC - A	COM	03783C100	2	12	SH		OTR	49	0	12	0
APPFOLIO INC - A	COM	03783C100	46	322	SH		DFND	49	322	0	0
APPHARVEST INC	COM	03783T103	2,047	111,869	SH		DFND	15	111,869	0	0
APPHARVEST INC	COM	03783T103	2,859	156,232	SH		DFND	22	156,232	0	0
APPHARVEST INC	COM	03783T103	617	33,723	SH		DFND	48	33,723	0	0
APPIAN CORP	CL A	03782L101	5,337	40,142	SH		DFND	15	28,277	0	11,865
APPIAN CORP	CL A	03782L101	12,188	91,671	SH		DFND	48	91,671	0	0
APPIAN CORP	CL A	03782L101	123	924	SH		DFND	49	924	0	0
APPLE HOSPITALITY REIT INC	COM	03784Y200	267	18,299	SH		OTR	15,119	18,299	0	0
APPLE HOSPITALITY REIT INC	COM	03784Y200	4,738	325,197	SH		DFND	15	228,468	0	96,729
APPLE HOSPITALITY REIT INC	COM	03784Y200	110	7,546	SH		OTR	22,87	7,546	0	0
APPLE HOSPITALITY REIT INC	COM	03784Y200	806	55,324	SH		OTR	22,78	55,324	0	0
APPLE HOSPITALITY REIT INC	COM	03784Y200	1,011	69,400	SH		DFND	22	69,400	0	0
APPLE HOSPITALITY REIT INC	COM	03784Y200	8,012	549,873	SH		DFND	48	487,385	0	62,488
APPLE HOSPITALITY REIT INC	COM	03784Y200	333	22,882	SH		DFND	49	22,789	0	93
APPLE HOSPITALITY REIT INC	COM	03784Y200	0	21	SH		DFND	59	21	0	0
APPLE INC	COM	037833100	173,051	1,416,705	SH		DFND	15,22	1,416,705	0	0
APPLE INC	COM	037833100	26,619	217,921	SH		OTR	15,119	217,921	0	0
APPLE INC	COM	037833100	65,165	533,485	SH		OTR	15,118	533,485	0	0
APPLE INC	COM	037833100	8,490,137	69,505,827	SH		DFND	15	42,720,776	0	26,785,051
APPLE INC	COM	037833100	779,596	6,382,284	SH		DFND	22	6,382,284	0	0
APPLE INC	COM	037833100	51,827	424,291	SH		DFND	4	423,419	0	872
APPLE INC	COM	037833100	26,710	218,667	SH		OTR	4	69,419	149,248	0
APPLE INC	COM	037833100	578,261	4,734,020	SH		DFND	40	4,269,315	0	464,705
APPLE INC	COM	037833100	6,528	53,441	SH		OTR	40,48	53,441	0	0
APPLE INC	COM	037833100	7,373	60,364	SH		DFND	42	60,364	0	0
APPLE INC	COM	037833100	6,350,066	51,985,802	SH		DFND	48	47,067,426	0	4,918,376
APPLE INC	COM	037833100	351,243	2,875,503	SH		OTR	49	1,128,786	1,263,379	483,338
APPLE INC	COM	037833100	1,941,343	15,893,107	SH		DFND	49	15,617,233	7,173	268,701
APPLE INC	COM	037833100	47,833	391,589	SH		DFND	59	374,773	0	16,816
APPLE INC	COM	037833100	89	726	SH		DFND		726	0	0
APPLE INC	COM	037833100	3,909	32,000	SH	Call	DFND	49	0	0	0
APPLIED GENETIC TECHNOLOGIES	COM	03820J100	114	22,515	SH		DFND	15	2,676	0	19,839
APPLIED GENETIC TECHNOLOGIES	COM	03820J100	354	69,887	SH		DFND	48	69,887	0	0
APPLIED INDUSTRIAL TECH INC	COM	03820C105	23,038	252,693	SH		DFND	15	226,899	0	25,794
APPLIED INDUSTRIAL TECH INC	COM	03820C105	137	1,507	SH		OTR	15,119	1,507	0	0
APPLIED INDUSTRIAL TECH INC	COM	03820C105	9,168	100,563	SH		DFND	22	100,563	0	0
APPLIED INDUSTRIAL TECH INC	COM	03820C105	59	645	SH		DFND	4	645	0	0
APPLIED INDUSTRIAL TECH INC	COM	03820C105	9,024	98,984	SH		DFND	48	98,984	0	0
APPLIED INDUSTRIAL TECH INC	COM	03820C105	3,652	40,053	SH		DFND	49	40,053	0	0
APPLIED MATERIALS INC	COM	038222105	731,145	5,472,643	SH		DFND	15	3,874,412	0	1,598,231
APPLIED MATERIALS INC	COM	038222105	1,756	13,144	SH		OTR	15,119	13,144	0	0
APPLIED MATERIALS INC	COM	038222105	11,032	82,577	SH		DFND	15,22	82,577	0	0
APPLIED MATERIALS INC	COM	038222105	15,142	113,337	SH		OTR	15,118	113,337	0	0
APPLIED MATERIALS INC	COM	038222105	210,543	1,575,919	SH		DFND	22	1,575,919	0	0
APPLIED MATERIALS INC	COM	038222105	129	966	SH		OTR	4	129	770	67
APPLIED MATERIALS INC	COM	038222105	1,400	10,479	SH		DFND	4	10,242	0	237
APPLIED MATERIALS INC	COM	038222105	420,168	3,144,971	SH		DFND	40	2,812,217	0	332,754
APPLIED MATERIALS INC	COM	038222105	3,838	28,728	SH		OTR	40,48	28,728	0	0
APPLIED MATERIALS INC	COM	038222105	4,218	31,575	SH		DFND	42	31,575	0	0
APPLIED MATERIALS INC	COM	038222105	385,472	2,885,270	SH		DFND	48	2,600,892	0	284,378
APPLIED MATERIALS INC	COM	038222105	68,105	509,770	SH		DFND	49	504,263	400	5,107
APPLIED MATERIALS INC	COM	038222105	8,566	64,114	SH		OTR	49	53,804	10,243	67
APPLIED MATERIALS INC	COM	038222105	3,982	29,805	SH		DFND	59	27,772	0	2,033
APPLIED MOLECULAR TRANSPORT	COM	03824M109	848	19,278	SH		DFND	15	11,913	0	7,365
APPLIED MOLECULAR TRANSPORT	COM	03824M109	2,092	47,543	SH		DFND	48	47,543	0	0
APPLIED OPTOELECTRONICS INC	COM	03823U102	955	114,188	SH		DFND	15	94,172	0	20,016
APPLIED OPTOELECTRONICS INC	COM	03823U102	479	57,293	SH		DFND	22	57,293	0	0
APPLIED OPTOELECTRONICS INC	COM	03823U102	461	55,138	SH		DFND	48	55,138	0	0
APPLIED OPTOELECTRONICS INC	COM	03823U102	15	1,755	SH		DFND	49	1,755	0	0
APPLIED THERAPEUTICS INC	COM	03828A101	375	19,998	SH		DFND	15	15,787	0	4,211
APPLIED THERAPEUTICS INC	COM	03828A101	757	40,372	SH		DFND	48	40,372	0	0
APREA THERAPEUTICS INC	COM	03836J102	77	15,016	SH		DFND	15	14,482	0	534
APREA THERAPEUTICS INC	COM	03836J102	164	32,187	SH		DFND	48	32,187	0	0
APREA THERAPEUTICS INC	COM	03836J102	3	580	SH		DFND	49	580	0	0
APRIA INC	COM	03836A101	7,447	266,614	SH		DFND	15	189,996	0	76,618
APRIA INC	COM	03836A101	6,769	242,368	SH		OTR	15,66	242,368	0	0
APRIA INC	COM	03836A101	19,481	697,511	SH		DFND	22	697,511	0	0
APRIA INC	COM	03836A101	571	20,460	SH		OTR	22,83	20,460	0	0
APRIA INC	COM	03836A101	2,260	80,906	SH		DFND	48	80,906	0	0
APTARGROUP INC	COM	038336103	279	1,969	SH		OTR	15,119	1,969	0	0
APTARGROUP INC	COM	038336103	55,598	392,447	SH		DFND	15	360,220	0	32,227
APTARGROUP INC	COM	038336103	10,882	76,810	SH		DFND	22	76,810	0	0
APTARGROUP INC	COM	038336103	153	1,077	SH		DFND	4	1,077	0	0
APTARGROUP INC	COM	038336103	34,416	242,934	SH		DFND	48	223,583	0	19,351
APTARGROUP INC	COM	038336103	432	3,052	SH		OTR	49	2,592	460	0
APTARGROUP INC	COM	038336103	5,966	42,109	SH		DFND	49	41,975	0	134
APTARGROUP INC	COM	038336103	15,362	108,436	SH		DFND	59	103,401	0	5,035
APTARGROUP INC	COM	038336103	0	1	SH		DFND		1	0	0
APTEVO THERAPEUTICS INC	COM	03835L207	157	5,130	SH		DFND	15	5,130	0	0
APTEVO THERAPEUTICS INC	COM	03835L207	148	4,834	SH		DFND	48	4,834	0	0
APTINYX INC	COM	03836N103	64	21,468	SH		DFND	15	2,815	0	18,653
APTINYX INC	COM	03836N103	305	101,652	SH		DFND	48	101,652	0	0
APTINYX INC	COM	03836N103	185	61,723	SH		DFND	49	61,723	0	0
APTIV PLC	SHS	G6095L109	1,255	9,104	SH		OTR	15,118	9,104	0	0
APTIV PLC	SHS	G6095L109	767	5,561	SH		OTR	15,119	5,561	0	0
APTIV PLC	SHS	G6095L109	3,313	24,026	SH		DFND	15,22	24,026	0	0
APTIV PLC	SHS	G6095L109	183,631	1,331,624	SH		DFND	15	848,699	0	482,925
APTIV PLC	SHS	G6095L109	8,038	58,292	SH		DFND	22	58,292	0	0
APTIV PLC	SHS	G6095L109	179	1,297	SH		DFND	4	1,297	0	0
APTIV PLC	SHS	G6095L109	108,087	783,809	SH		DFND	48	703,042	0	80,767
APTIV PLC	SHS	G6095L109	7,096	51,455	SH		DFND	49	51,376	0	79
APTIV PLC	SHS	G6095L109	844	6,123	SH		OTR	49	4,575	1,548	0
APTIV PLC	SHS	G6095L109	356	2,580	SH		DFND	59	2,485	0	95
APYX MEDICAL CORP	COM	03837C106	238	24,679	SH		DFND	15	5,153	0	19,526
APYX MEDICAL CORP	COM	03837C106	812	84,038	SH		DFND	48	84,038	0	0
AQUA METALS INC	COM	03837J101	124	30,608	SH		DFND	48	30,608	0	0
AQUABOUNTY TECHNOLOGIES	COM	03842K200	210	31,283	SH		DFND	48	31,283	0	0
AQUABOUNTY TECHNOLOGIES	COM	03842K200	0	7	SH		DFND		7	0	0
AQUESTIVE THERAPEUTICS INC	COM	03843E104	538	103,488	SH		DFND	15	102,048	0	1,440
AQUESTIVE THERAPEUTICS INC	COM	03843E104	316	60,746	SH		DFND	48	60,746	0	0
AQUESTIVE THERAPEUTICS INC	COM	03843E104	0	25	SH		DFND	49	25	0	0
ARAMARK	COM	03852U106	22,588	597,873	SH		DFND	15,22	597,873	0	0
ARAMARK	COM	03852U106	41,092	1,087,670	SH		DFND	15	661,196	0	426,474
ARAMARK	COM	03852U106	507	13,408	SH		OTR	15,119	13,408	0	0
ARAMARK	COM	03852U106	21,486	568,724	SH		DFND	22	568,724	0	0
ARAMARK	COM	03852U106	8	202	SH		OTR	4	0	0	202
ARAMARK	COM	03852U106	10	265	SH		DFND	4	265	0	0
ARAMARK	COM	03852U106	558	14,777	SH		DFND	42	14,777	0	0
ARAMARK	COM	03852U106	38,335	1,014,683	SH		DFND	48	944,831	0	69,852
ARAMARK	COM	03852U106	166	4,399	SH		DFND	49	4,399	0	0
ARAVIVE INC	COM	03890D108	15	2,302	SH		DFND	15	1,394	0	908
ARAVIVE INC	COM	03890D108	263	39,945	SH		DFND	48	39,945	0	0
ARAVIVE INC	COM	03890D108	2	340	SH		DFND	49	340	0	0
ARBOR REALTY TRUST INC	COM	038923108	2,596	163,261	SH		DFND	15	59,395	0	103,866
ARBOR REALTY TRUST INC	COM	038923108	4,557	286,609	SH		DFND	48	286,609	0	0
ARBOR REALTY TRUST INC	COM	038923108	29	1,820	SH		DFND	49	1,820	0	0
ARBOR REALTY TRUST INC	COM	038923108	330	20,726	SH		DFND	59	20,726	0	0
ARBOR REALTY TRUST INC	COM	038923108	0	1	SH		DFND		1	0	0
ARBUTUS BIOPHARMA CORP	COM	03879J100	171	51,416	SH		DFND	48	51,416	0	0
ARC DOCUMENT SOLUTIONS INC	COM	00191G103	63	29,637	SH		DFND	48	29,637	0	0
ARCBEST CORP	COM	03937C105	12,494	177,546	SH		DFND	15	155,864	0	21,682
ARCBEST CORP	COM	03937C105	4,588	65,196	SH		DFND	22	65,196	0	0
ARCBEST CORP	COM	03937C105	4,510	64,085	SH		DFND	48	64,085	0	0
ARCBEST CORP	COM	03937C105	287	4,085	SH		DFND	49	4,085	0	0
ARCBEST CORP	COM	03937C105	3	45	SH		OTR	49	0	45	0
ARCBEST CORP	COM	03937C105	0	1	SH		DFND		1	0	0
ARCELORMITTAL-NY REGISTERED	NY R	03938L203	20	675	SH		DFND	4	675	0	0
ARCELORMITTAL-NY REGISTERED	NY R	03938L203	5	181	SH		OTR	49	0	181	0
ARCELORMITTAL-NY REGISTERED	NY R	03938L203	255	8,735	SH		DFND	49	8,735	0	0
ARCELORMITTAL-NY REGISTERED	NY R	03938L203	0	3	SH		DFND	59	3	0	0
ARCH CAPITAL GROUP LTD	ORD	G0450A105	29,970	781,090	SH		DFND	15	442,340	0	338,750
ARCH CAPITAL GROUP LTD	ORD	G0450A105	685	17,858	SH		OTR	15,119	17,858	0	0
ARCH CAPITAL GROUP LTD	ORD	G0450A105	14,846	386,927	SH		DFND	22	386,927	0	0
ARCH CAPITAL GROUP LTD	ORD	G0450A105	25,958	676,519	SH		DFND	48	558,848	0	117,671
ARCH CAPITAL GROUP LTD	ORD	G0450A105	1,230	32,058	SH		DFND	49	32,058	0	0
ARCH CAPITAL GROUP LTD	ORD	G0450A105	626	16,326	SH		OTR	49	0	16,326	0
ARCH CAPITAL GROUP LTD	ORD	G0450A105	223	5,808	SH		DFND	59	5,808	0	0
ARCH RESOURCES INC	CL A	03940R107	1,469	35,320	SH		DFND	15	26,018	0	9,302
ARCH RESOURCES INC	CL A	03940R107	1,569	37,705	SH		DFND	48	37,705	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	1,777	31,176	SH		OTR	15,119	31,176	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	2,858	50,135	SH		DFND	15,22	50,135	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	1,073	18,820	SH		OTR	15,118	18,820	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	224,836	3,944,493	SH		DFND	15	3,054,575	0	889,918
ARCHER-DANIELS-MIDLAND CO	COM	039483102	6,561	115,108	SH		DFND	22	115,108	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	236	4,141	SH		DFND	4	4,141	0	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	26	455	SH		OTR	4	0	455	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	93,004	1,631,646	SH		DFND	48	1,463,334	0	168,312
ARCHER-DANIELS-MIDLAND CO	COM	039483102	692	12,134	SH		OTR	49	8,247	3,887	0
ARCHER-DANIELS-MIDLAND CO	COM	039483102	9,817	172,220	SH		DFND	49	172,005	0	215
ARCHER-DANIELS-MIDLAND CO	COM	039483102	3,490	61,223	SH		DFND	59	60,385	0	838
ARCHROCK INC	COM	03957W106	22,715	2,393,611	SH		DFND	15	2,280,409	0	113,202
ARCHROCK INC	COM	03957W106	3,211	338,402	SH		DFND	22	338,402	0	0
ARCHROCK INC	COM	03957W106	22	2,290	SH		DFND	4	2,290	0	0
ARCHROCK INC	COM	03957W106	3,038	320,081	SH		DFND	48	320,081	0	0
ARCHROCK INC	COM	03957W106	1,183	124,620	SH		DFND	49	124,620	0	0
ARCHROCK INC	COM	03957W106	911	95,967	SH		DFND	59	93,580	0	2,387
ARCIMOTO INC	COM	039587100	267	20,177	SH		DFND	48	20,177	0	0
ARCIMOTO INC	COM	039587100	165	12,450	SH		OTR	49	12,450	0	0
ARCIMOTO INC	COM	039587100	1	100	SH		DFND	49	100	0	0
ARCO PLATFORM LTD - CLASS A	COM	G04553106	1,041	41,078	SH		OTR	40,48	41,078	0	0
ARCO PLATFORM LTD - CLASS A	COM	G04553106	3,916	154,533	SH		DFND	40	148,723	0	5,810
ARCO PLATFORM LTD - CLASS A	COM	G04553106	1,949	76,928	SH		DFND	40,22	76,928	0	0
ARCONIC CORP	COM	03966V107	250	9,862	SH		OTR	15,119	9,862	0	0
ARCONIC CORP	COM	03966V107	12,764	502,714	SH		DFND	15	414,708	0	88,006
ARCONIC CORP	COM	03966V107	6,359	250,447	SH		DFND	22	250,447	0	0
ARCONIC CORP	COM	03966V107	6	247	SH		DFND	4	247	0	0
ARCONIC CORP	COM	03966V107	5	205	SH		OTR	4	24	0	181
ARCONIC CORP	COM	03966V107	6,302	248,204	SH		DFND	48	248,204	0	0
ARCONIC CORP	COM	03966V107	197	7,769	SH		DFND	49	7,767	0	2
ARCONIC CORP	COM	03966V107	389	15,327	SH		OTR	49	1,313	13,148	866
ARCONIC CORP	COM	03966V107	0	1	SH		DFND		1	0	0
ARCOSA INC	COM	039653100	18,085	277,842	SH		DFND	15	234,224	0	43,618
ARCOSA INC	COM	039653100	8,089	124,272	SH		DFND	22	124,272	0	0
ARCOSA INC	COM	039653100	63	963	SH		DFND	4	963	0	0
ARCOSA INC	COM	039653100	8,017	123,169	SH		DFND	48	123,169	0	0
ARCOSA INC	COM	039653100	2,109	32,408	SH		DFND	49	32,408	0	0
ARCOSA INC	COM	039653100	177	2,717	SH		OTR	49	14	37	2,666
ARCOSA INC	COM	039653100	0	1	SH		DFND		1	0	0
ARCTURUS THERAPEUTICS HOLDIN	COM	03969T109	1,041	25,217	SH		DFND	15	18,464	0	6,753
ARCTURUS THERAPEUTICS HOLDIN	COM	03969T109	2,004	48,518	SH		DFND	48	48,518	0	0
ARCTURUS THERAPEUTICS HOLDIN	COM	03969T109	2	50	SH		DFND	49	50	0	0
ARCUS BIOSCIENCES INC	COM	03969F109	1,495	53,235	SH		DFND	15	36,351	0	16,884
ARCUS BIOSCIENCES INC	COM	03969F109	3,020	107,549	SH		DFND	48	107,549	0	0
ARCUS BIOSCIENCES INC	COM	03969F109	252	8,965	SH		DFND	49	8,965	0	0
ARCUTIS BIOTHERAPEUTICS INC	COM	03969K108	841	29,080	SH		DFND	15	23,236	0	5,844
ARCUTIS BIOTHERAPEUTICS INC	COM	03969K108	1,673	57,821	SH		DFND	48	57,821	0	0
ARDAGH GROUP SA	CL A	L0223L101	547	21,546	SH		DFND	15	12,867	0	8,679
ARDAGH GROUP SA	CL A	L0223L101	557	21,908	SH		DFND	48	16,637	0	5,271
ARDAGH GROUP SA	CL A	L0223L101	14	542	SH		DFND	49	542	0	0
ARDAGH GROUP SA	CL A	L0223L101	0	12	SH		DFND		12	0	0
ARDELYX INC	COM	039697107	659	99,488	SH		DFND	15	71,617	0	27,871
ARDELYX INC	COM	039697107	1,244	187,857	SH		DFND	48	187,857	0	0
ARDMORE SHIPPING CORP	COM	Y0207T100	103	22,593	SH		DFND	15	3,494	0	19,099
ARDMORE SHIPPING CORP	COM	Y0207T100	336	73,998	SH		DFND	48	73,998	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	15,002	216,196	SH		DFND	15	158,113	0	58,083
ARENA PHARMACEUTICALS INC	COM	040047607	20,860	300,626	SH		DFND	15,22	300,626	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	1,167	16,824	SH		OTR	22,78	16,824	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	942	13,577	SH		OTR	22,87	13,577	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	13,856	199,677	SH		DFND	22	199,677	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	500	7,210	SH		DFND	42	7,210	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	12,186	175,622	SH		DFND	48	175,622	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	54	783	SH		DFND	49	783	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	491	7,075	SH		DFND	59	7,075	0	0
ARENA PHARMACEUTICALS INC	COM	040047607	0	1	SH		DFND		1	0	0
ARES CAPITAL CORP	COM	04010L103	6,082	325,055	SH		DFND	15	325,055	0	0
ARES CAPITAL CORP	COM	04010L103	53	2,845	SH		OTR	4	2,845	0	0
ARES CAPITAL CORP	COM	04010L103	401	21,438	SH		DFND	4	21,438	0	0
ARES CAPITAL CORP	COM	04010L103	1,343	71,784	SH		OTR	49	45,842	25,942	0
ARES CAPITAL CORP	COM	04010L103	20,133	1,076,061	SH		DFND	49	1,065,307	0	10,754
ARES CAPITAL CORP	COM	04010L103	2	86	SH		DFND	59	86	0	0
ARES CAPITAL CORP	COM	04010L103	0	1	SH		DFND		1	0	0
ARES CAPITAL CORP	NOTE	04010LAT0	258	250,000	SH		DFND	15	250,000	0	0
ARES CAPITAL CORP	NOTE	04010LAW3	544	500,000	SH		DFND	15	500,000	0	0
ARES COMMERCIAL REAL ESTATE	COM	04013V108	944	68,791	SH		DFND	15	45,908	0	22,883
ARES COMMERCIAL REAL ESTATE	COM	04013V108	1,164	84,840	SH		DFND	48	84,840	0	0
ARES COMMERCIAL REAL ESTATE	COM	04013V108	32	2,346	SH		DFND	49	2,346	0	0
ARES COMMERCIAL REAL ESTATE	COM	04013V108	0	1	SH		DFND		1	0	0
ARES MANAGEMENT CORP - A	CL A	03990B101	49,600	885,241	SH		DFND	15	718,357	0	166,884
ARES MANAGEMENT CORP - A	CL A	03990B101	42,976	767,020	SH		DFND	15,22	767,020	0	0
ARES MANAGEMENT CORP - A	CL A	03990B101	17,616	314,397	SH		DFND	22	314,397	0	0
ARES MANAGEMENT CORP - A	CL A	03990B101	3	60	SH		OTR	4	0	0	60
ARES MANAGEMENT CORP - A	CL A	03990B101	1,075	19,186	SH		DFND	42	19,186	0	0
ARES MANAGEMENT CORP - A	CL A	03990B101	11,872	211,892	SH		DFND	48	182,935	0	28,957
ARES MANAGEMENT CORP - A	CL A	03990B101	53	946	SH		DFND	49	946	0	0
ARES MANAGEMENT CORP - A	CL A	03990B101	12	221	SH		OTR	49	0	221	0
ARES MANAGEMENT CORP - A	CL A	03990B101	0	1	SH		DFND		1	0	0
ARGAN INC	COM	04010E109	3,754	70,373	SH		DFND	15	56,803	0	13,570
ARGAN INC	COM	04010E109	2,003	37,541	SH		DFND	48	37,541	0	0
ARGAN INC	COM	04010E109	5	100	SH		DFND	49	100	0	0
ARGENX SE - ADR	SPON	04016X101	685	2,487	SH		OTR	22,78	2,487	0	0
ARGENX SE - ADR	SPON	04016X101	155	564	SH		DFND	49	564	0	0
ARGENX SE - ADR	SPON	04016X101	335	1,217	SH		OTR	49	1,187	30	0
ARGENX SE - ADR	SPON	04016X101	0	1	SH		DFND		1	0	0
ARGO GROUP INTERNATIONAL	COM	G0464B107	2,382	47,343	SH		DFND	15	18,172	0	29,171
ARGO GROUP INTERNATIONAL	COM	G0464B107	4,459	88,612	SH		DFND	48	88,612	0	0
ARGO GROUP INTERNATIONAL	COM	G0464B107	411	8,164	SH		DFND	49	8,164	0	0
ARGO GROUP INTERNATIONAL	COM	G0464B107	16	310	SH		DFND	59	310	0	0
ARISTA NETWORKS INC	COM	040413106	193	640	SH		OTR	15,119	640	0	0
ARISTA NETWORKS INC	COM	040413106	558	1,847	SH		OTR	15,118	1,847	0	0
ARISTA NETWORKS INC	COM	040413106	1,501	4,972	SH		DFND	15,22	4,972	0	0
ARISTA NETWORKS INC	COM	040413106	62,016	205,426	SH		DFND	15	114,570	0	90,856
ARISTA NETWORKS INC	COM	040413106	4,576	15,158	SH		DFND	22	15,158	0	0
ARISTA NETWORKS INC	COM	040413106	87	289	SH		DFND	4	289	0	0
ARISTA NETWORKS INC	COM	040413106	49,140	162,773	SH		DFND	48	144,566	0	18,207
ARISTA NETWORKS INC	COM	040413106	3,187	10,557	SH		OTR	49	2,564	7,993	0
ARISTA NETWORKS INC	COM	040413106	3,080	10,201	SH		DFND	49	10,161	0	40
ARISTA NETWORKS INC	COM	040413106	302	1,001	SH		DFND	59	1,001	0	0
ARK AUTONOMOUS TECH & ROBOT	AUTN	00214Q203	1,257	14,970	SH		OTR	49	14,970	0	0
ARK AUTONOMOUS TECH & ROBOT	AUTN	00214Q203	84	1,000	SH		DFND	49	1,000	0	0
ARK GENOMIC REVOLUTION ETF	GENO	00214Q302	211	2,379	SH		DFND	49	2,379	0	0
ARK GENOMIC REVOLUTION ETF	GENO	00214Q302	399	4,498	SH		OTR	49	4,013	485	0
ARK INNOVATION ETF	INNO	00214Q104	682	5,684	SH		DFND	49	5,584	0	100
ARK INNOVATION ETF	INNO	00214Q104	82	680	SH		OTR	49	480	200	0
ARKO CORP	COM	041242108	182	18,300	SH		DFND	15	18,300	0	0
ARKO CORP	COM	041242108	247	24,836	SH		DFND	48	24,836	0	0
ARLINGTON ASSET INVESTMENT-A	CL A	041356205	170	42,180	SH		DFND	15	18,532	0	23,648
ARLINGTON ASSET INVESTMENT-A	CL A	041356205	326	80,793	SH		DFND	48	80,793	0	0
ARLINGTON ASSET INVESTMENT-A	CL A	041356205	7	1,725	SH		DFND	49	1,725	0	0
ARLO TECHNOLOGIES INC	COM	04206A101	2,344	373,320	SH		DFND	15	300,699	0	72,621
ARLO TECHNOLOGIES INC	COM	04206A101	5,883	936,776	SH		DFND	22	936,776	0	0
ARLO TECHNOLOGIES INC	COM	04206A101	2,201	350,459	SH		DFND	48	350,459	0	0
ARLO TECHNOLOGIES INC	COM	04206A101	51	8,195	SH		DFND	49	8,195	0	0
ARMADA HOFFLER PROPERTIES IN	COM	04208T108	5,509	439,305	SH		DFND	15	388,718	0	50,587
ARMADA HOFFLER PROPERTIES IN	COM	04208T108	1,929	153,855	SH		DFND	22	153,855	0	0
ARMADA HOFFLER PROPERTIES IN	COM	04208T108	14	1,095	SH		DFND	4	1,095	0	0
ARMADA HOFFLER PROPERTIES IN	COM	04208T108	1,808	144,203	SH		DFND	48	144,203	0	0
ARMADA HOFFLER PROPERTIES IN	COM	04208T108	1,022	81,486	SH		DFND	49	81,486	0	0
ARMOUR RESIDENTIAL REIT INC	COM	042315507	3,687	302,212	SH		DFND	15	246,880	0	55,332
ARMOUR RESIDENTIAL REIT INC	COM	042315507	2,032	166,565	SH		DFND	22	166,565	0	0
ARMOUR RESIDENTIAL REIT INC	COM	042315507	1,950	159,843	SH		DFND	48	159,843	0	0
ARMOUR RESIDENTIAL REIT INC	COM	042315507	159	13,052	SH		DFND	49	13,052	0	0
ARMOUR RESIDENTIAL REIT INC	COM	042315507	0	3	SH		DFND	59	3	0	0
ARMOUR RESIDENTIAL REIT INC	COM	042315507	0	1	SH		DFND		1	0	0
ARMSTRONG FLOORING INC	COM	04238R106	192	39,224	SH		DFND	48	39,224	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	30,344	336,821	SH		DFND	15	266,281	0	70,540
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	23,347	259,152	SH		DFND	15,22	259,152	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	6,530	72,486	SH		OTR	15,66	72,486	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	11,477	127,393	SH		DFND	22	127,393	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	1,994	22,130	SH		OTR	22,87	22,130	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	577	6,409	SH		DFND	42	6,409	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	18,744	208,064	SH		DFND	48	193,042	0	15,022
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	58	647	SH		DFND	49	617	0	30
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	86	960	SH		OTR	49	0	960	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	0	1	SH		DFND	59	1	0	0
ARMSTRONG WORLD INDUSTRIES	COM	04247X102	0	1	SH		DFND		1	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	34,685	1,163,144	SH		DFND	15	1,028,377	0	134,767
ARRAY TECHNOLOGIES INC	COM	04271T100	31,449	1,054,621	SH		DFND	15,22	1,054,621	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	27,552	923,954	SH		DFND	22	923,954	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	471	15,800	SH		OTR	22,104	15,800	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	63	2,110	SH		DFND	4	2,110	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	1	50	SH		OTR	4	50	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	768	25,771	SH		DFND	42	25,771	0	0
ARRAY TECHNOLOGIES INC	COM	04271T100	9,487	318,152	SH		DFND	48	281,059	0	37,093
ARRAY TECHNOLOGIES INC	COM	04271T100	2,527	84,757	SH		DFND	49	83,747	0	1,010
ARRAY TECHNOLOGIES INC	COM	04271T100	384	12,870	SH		OTR	49	10,668	2,162	40
ARROW ELECTRONICS INC	COM	042735100	46,854	422,797	SH		DFND	15	380,663	0	42,134
ARROW ELECTRONICS INC	COM	042735100	676	6,096	SH		OTR	15,119	6,096	0	0
ARROW ELECTRONICS INC	COM	042735100	392	3,540	SH		DFND	15,22	3,540	0	0
ARROW ELECTRONICS INC	COM	042735100	9,445	85,226	SH		DFND	22	85,226	0	0
ARROW ELECTRONICS INC	COM	042735100	125	1,130	SH		DFND	4	1,130	0	0
ARROW ELECTRONICS INC	COM	042735100	7	66	SH		OTR	4	0	66	0
ARROW ELECTRONICS INC	COM	042735100	31,949	288,298	SH		DFND	48	265,094	0	23,204
ARROW ELECTRONICS INC	COM	042735100	91	824	SH		OTR	49	4	820	0
ARROW ELECTRONICS INC	COM	042735100	4,939	44,572	SH		DFND	49	44,422	0	150
ARROW ELECTRONICS INC	COM	042735100	927	8,369	SH		DFND	59	8,369	0	0
ARROW ELECTRONICS INC	COM	042735100	0	1	SH		DFND		1	0	0
ARROW FINANCIAL CORP	COM	042744102	2,591	77,774	SH		DFND	15	63,765	0	14,009
ARROW FINANCIAL CORP	COM	042744102	1,186	35,601	SH		DFND	48	35,601	0	0
ARROW FINANCIAL CORP	COM	042744102	12	369	SH		DFND	49	369	0	0
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	26,122	393,943	SH		DFND	15	359,583	0	34,360
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	55	827	SH		OTR	15,119	827	0	0
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	7,316	110,325	SH		DFND	22	110,325	0	0
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	102	1,533	SH		DFND	4	1,533	0	0
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	30,904	466,050	SH		DFND	48	466,050	0	0
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	2,844	42,887	SH		DFND	49	42,687	0	200
ARROWHEAD PHARMACEUTICALS IN	COM	04280A100	5	71	SH		OTR	49	0	71	0
ARTELO BIOSCIENCES INC	COM	04301G201	167	102,390	SH		DFND	48	102,390	0	0
ARTESIAN RESOURCES CORP-CL A	CL A	043113208	3,298	83,752	SH		DFND	15	78,238	0	5,514
ARTESIAN RESOURCES CORP-CL A	CL A	043113208	972	24,690	SH		DFND	48	24,690	0	0
ARTESIAN RESOURCES CORP-CL A	CL A	043113208	10	250	SH		DFND	49	250	0	0
ARTESIAN RESOURCES CORP-CL A	CL A	043113208	0	1	SH		DFND		1	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	598	4,792	SH		OTR	15,119	4,792	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	829	6,645	SH		OTR	15,118	6,645	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	96,942	776,969	SH		DFND	15	467,216	0	309,753
ARTHUR J GALLAGHER & CO	COM	363576109	2,152	17,245	SH		DFND	15,22	17,245	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	3,837	30,754	SH		DFND	22	30,754	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	668	5,357	SH		DFND	4	5,357	0	0
ARTHUR J GALLAGHER & CO	COM	363576109	64	509	SH		OTR	4	37	450	22
ARTHUR J GALLAGHER & CO	COM	363576109	69,883	560,094	SH		DFND	48	502,643	0	57,451
ARTHUR J GALLAGHER & CO	COM	363576109	1,701	13,637	SH		OTR	49	7,931	5,706	0
ARTHUR J GALLAGHER & CO	COM	363576109	30,506	244,496	SH		DFND	49	242,337	0	2,159
ARTHUR J GALLAGHER & CO	COM	363576109	22,875	183,334	SH		DFND	59	174,575	0	8,759
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	27,312	523,524	SH		DFND	15	485,975	0	37,549
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	1,583	30,352	SH		OTR	22,121	30,352	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	3,963	75,970	SH		OTR	22,96	75,970	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	7,325	140,397	SH		DFND	48	140,397	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	53	1,009	SH		DFND	49	1,009	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	16	316	SH		OTR	49	316	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	256	4,905	SH		DFND	59	4,905	0	0
ARTISAN PARTNERS ASSET MA -A	CL A	04316A108	0	1	SH		DFND		1	0	0
ART'S-WAY MANUFACTURING CO	COM	043168103	36	11,689	SH		DFND	48	11,689	0	0
ART'S-WAY MANUFACTURING CO	COM	043168103	0	1	SH		DFND		1	0	0
ARVINAS INC	COM	04335A105	2,754	41,667	SH		DFND	15	29,234	0	12,433
ARVINAS INC	COM	04335A105	5,682	85,964	SH		DFND	48	85,964	0	0
ASANA INC - CL A	CL A	04342Y104	16,961	593,458	SH		DFND	15	593,458	0	0
ASANA INC - CL A	CL A	04342Y104	147	5,127	SH		DFND	48	5,127	0	0
ASANA INC - CL A	CL A	04342Y104	662	23,175	SH		DFND	49	23,175	0	0
ASBURY AUTOMOTIVE GROUP	COM	043436104	299	1,522	SH		OTR	15,119	1,522	0	0
ASBURY AUTOMOTIVE GROUP	COM	043436104	21,642	110,135	SH		DFND	15	97,272	0	12,863
ASBURY AUTOMOTIVE GROUP	COM	043436104	9,759	49,662	SH		DFND	22	49,662	0	0
ASBURY AUTOMOTIVE GROUP	COM	043436104	3	16	SH		DFND	4	16	0	0
ASBURY AUTOMOTIVE GROUP	COM	043436104	10,253	52,178	SH		DFND	48	52,178	0	0
ASBURY AUTOMOTIVE GROUP	COM	043436104	1,157	5,887	SH		DFND	49	5,887	0	0
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	26,101	202,518	SH		DFND	15,22	202,518	0	0
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	4,875	37,824	SH		DFND	15	32,071	0	5,753
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	1,084	8,413	SH		DFND	22	8,413	0	0
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	575	4,463	SH		DFND	42	4,463	0	0
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	1,170	9,078	SH		DFND	48	9,078	0	0
ASCENDIS PHARMA A/S - ADR	SPON	04351P101	67	518	SH		OTR	49	449	69	0
ASENSUS SURGICAL INC	COM	04367G103	227	69,766	SH		DFND	48	69,766	0	0
ASENSUS SURGICAL INC	COM	04367G103	0	1	SH		DFND		1	0	0
ASGN INC	COM	00191U102	18,694	195,871	SH		DFND	15	174,098	0	21,773
ASGN INC	COM	00191U102	420	4,400	SH		DFND	15,22	4,400	0	0
ASGN INC	COM	00191U102	202	2,112	SH		OTR	15,119	2,112	0	0
ASGN INC	COM	00191U102	4,032	42,249	SH		OTR	22,80	42,249	0	0
ASGN INC	COM	00191U102	6,016	63,039	SH		DFND	22	63,039	0	0
ASGN INC	COM	00191U102	227	2,376	SH		OTR	22,87	2,376	0	0
ASGN INC	COM	00191U102	812	8,510	SH		OTR	22,104	8,510	0	0
ASGN INC	COM	00191U102	21	216	SH		OTR	4	0	0	216
ASGN INC	COM	00191U102	37	385	SH		DFND	4	385	0	0
ASGN INC	COM	00191U102	22,273	233,367	SH		DFND	48	233,367	0	0
ASGN INC	COM	00191U102	1,487	15,584	SH		DFND	49	15,484	0	100
ASGN INC	COM	00191U102	1,826	19,130	SH		OTR	49	19,054	76	0
ASGN INC	COM	00191U102	204	2,137	SH		DFND	59	2,042	0	95
ASGN INC	COM	00191U102	0	1	SH		DFND		1	0	0
ASHFORD HOSPITALITY TRUST	COM	044103877	268	90,982	SH		DFND	48	90,982	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	235	2,650	SH		OTR	15,119	2,650	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	492	5,540	SH		DFND	15,22	5,540	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	24,230	272,953	SH		DFND	15	248,874	0	24,079
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	9,276	104,495	SH		DFND	22	104,495	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	22	245	SH		DFND	4	245	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	1,394	15,700	SH		OTR	4	0	15,700	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	18,647	210,060	SH		DFND	48	192,156	0	17,904
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	15	173	SH		OTR	49	85	88	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	2,822	31,786	SH		DFND	49	31,686	0	100
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	796	8,967	SH		DFND	59	8,967	0	0
ASHLAND GLOBAL HOLDINGS INC	COM	044186104	0	1	SH		DFND		1	0	0
ASML HOLDING NV-NY REG SHS	N Y	N07059210	52,529	85,087	SH		DFND	15	85,087	0	0
ASML HOLDING NV-NY REG SHS	N Y	N07059210	1,532	2,481	SH		OTR	4	76	2,200	205
ASML HOLDING NV-NY REG SHS	N Y	N07059210	549	889	SH		DFND	4	889	0	0
ASML HOLDING NV-NY REG SHS	N Y	N07059210	24,684	39,983	SH		DFND	42	39,983	0	0
ASML HOLDING NV-NY REG SHS	N Y	N07059210	22,053	35,721	SH		DFND	49	35,486	0	235
ASML HOLDING NV-NY REG SHS	N Y	N07059210	1,293	2,095	SH		OTR	49	1,040	1,055	0
ASML HOLDING NV-NY REG SHS	N Y	N07059210	6,458	10,461	SH		DFND	59	9,915	0	546
ASPEN AEROGELS INC	COM	04523Y105	489	24,065	SH		DFND	15	2,903	0	21,162
ASPEN AEROGELS INC	COM	04523Y105	1,076	52,888	SH		DFND	48	52,888	0	0
ASPEN AEROGELS INC	COM	04523Y105	6	307	SH		DFND	49	307	0	0
ASPEN GROUP INC	COM	04530L203	21	3,530	SH		DFND	15	2,127	0	1,403
ASPEN GROUP INC	COM	04530L203	313	52,152	SH		DFND	48	52,152	0	0
ASPEN TECHNOLOGY INC	COM	045327103	38,577	267,281	SH		DFND	15	235,859	0	31,422
ASPEN TECHNOLOGY INC	COM	045327103	780	5,404	SH		DFND	22	5,404	0	0
ASPEN TECHNOLOGY INC	COM	045327103	17,966	124,481	SH		DFND	48	103,376	0	21,105
ASPEN TECHNOLOGY INC	COM	045327103	192	1,331	SH		DFND	49	1,331	0	0
ASPEN TECHNOLOGY INC	COM	045327103	12	83	SH		OTR	49	0	83	0
ASPEN TECHNOLOGY INC	COM	045327103	494	3,426	SH		DFND	59	3,284	0	142
ASPIRA WOMEN'S HEALTH INC	COM	04537Y109	319	47,199	SH		DFND	15	11,723	0	35,476
ASPIRA WOMEN'S HEALTH INC	COM	04537Y109	1,207	178,752	SH		DFND	48	178,752	0	0
ASSEMBLY BIOSCIENCES INC	COM	045396108	231	50,245	SH		DFND	15	30,396	0	19,849
ASSEMBLY BIOSCIENCES INC	COM	045396108	352	76,420	SH		DFND	48	76,420	0	0
ASSERTIO HOLDINGS INC	COM	04546C106	28	41,552	SH		DFND	48	41,552	0	0
ASSETMARK FINANCIAL HOLDINGS	COM	04546L106	494	21,166	SH		DFND	15	10,615	0	10,551
ASSETMARK FINANCIAL HOLDINGS	COM	04546L106	1,010	43,256	SH		DFND	48	43,256	0	0
ASSOCIATED BANC-CORP	COM	045487105	25,540	1,196,808	SH		DFND	15	1,122,857	0	73,951
ASSOCIATED BANC-CORP	COM	045487105	1,085	50,830	SH		DFND	15,22	50,830	0	0
ASSOCIATED BANC-CORP	COM	045487105	246	11,509	SH		OTR	15,119	11,509	0	0
ASSOCIATED BANC-CORP	COM	045487105	2,148	100,653	SH		OTR	22,121	100,653	0	0
ASSOCIATED BANC-CORP	COM	045487105	11,896	557,461	SH		DFND	22	557,461	0	0
ASSOCIATED BANC-CORP	COM	045487105	108	5,046	SH		DFND	4	5,046	0	0
ASSOCIATED BANC-CORP	COM	045487105	12,186	571,023	SH		DFND	48	523,287	0	47,736
ASSOCIATED BANC-CORP	COM	045487105	10	475	SH		OTR	49	60	415	0
ASSOCIATED BANC-CORP	COM	045487105	3,751	175,781	SH		DFND	49	175,054	0	727
ASSOCIATED CAPITAL GROUP - A	CL A	045528106	131	3,651	SH		DFND	15	1,057	0	2,594
ASSOCIATED CAPITAL GROUP - A	CL A	045528106	169	4,711	SH		DFND	48	4,711	0	0
ASSOCIATED CAPITAL GROUP - A	CL A	045528106	20	550	SH		DFND	49	550	0	0
ASSOCIATED CAPITAL GROUP - A	CL A	045528106	0	1	SH		DFND		1	0	0
ASSURANT INC	COM	04621X108	11,196	78,973	SH		OTR	15,118	78,973	0	0
ASSURANT INC	COM	04621X108	171,363	1,208,737	SH		DFND	15	1,057,519	0	151,218
ASSURANT INC	COM	04621X108	755	5,323	SH		DFND	15,22	5,323	0	0
ASSURANT INC	COM	04621X108	501	3,537	SH		OTR	15,119	3,537	0	0
ASSURANT INC	COM	04621X108	54,610	385,198	SH		DFND	22	385,198	0	0
ASSURANT INC	COM	04621X108	282	1,990	SH		DFND	4	1,990	0	0
ASSURANT INC	COM	04621X108	24	166	SH		OTR	4	36	130	0
ASSURANT INC	COM	04621X108	270	1,901	SH		DFND	42	1,901	0	0
ASSURANT INC	COM	04621X108	36,126	254,824	SH		DFND	48	236,485	0	18,339
ASSURANT INC	COM	04621X108	1,044	7,363	SH		OTR	49	5,194	2,138	31
ASSURANT INC	COM	04621X108	8,716	61,477	SH		DFND	49	60,814	0	663
ASSURED GUARANTY LTD	COM	G0585R106	323	7,642	SH		OTR	15,119	7,642	0	0
ASSURED GUARANTY LTD	COM	G0585R106	14,033	331,895	SH		DFND	15	296,546	0	35,349
ASSURED GUARANTY LTD	COM	G0585R106	9,333	220,738	SH		DFND	22	220,738	0	0
ASSURED GUARANTY LTD	COM	G0585R106	58	1,375	SH		DFND	4	1,375	0	0
ASSURED GUARANTY LTD	COM	G0585R106	6,468	152,980	SH		DFND	48	129,222	0	23,758
ASSURED GUARANTY LTD	COM	G0585R106	1,301	30,779	SH		DFND	49	30,779	0	0
ASTEC INDUSTRIES INC	COM	046224101	9,565	126,817	SH		OTR	15,66	126,817	0	0
ASTEC INDUSTRIES INC	COM	046224101	19,625	260,213	SH		DFND	15	200,992	0	59,221
ASTEC INDUSTRIES INC	COM	046224101	7,099	94,129	SH		DFND	22	94,129	0	0
ASTEC INDUSTRIES INC	COM	046224101	1,577	20,916	SH		OTR	22,121	20,916	0	0
ASTEC INDUSTRIES INC	COM	046224101	2,797	37,083	SH		OTR	22,114	37,083	0	0
ASTEC INDUSTRIES INC	COM	046224101	16	215	SH		DFND	4	215	0	0
ASTEC INDUSTRIES INC	COM	046224101	4,519	59,921	SH		DFND	48	59,921	0	0
ASTEC INDUSTRIES INC	COM	046224101	612	8,115	SH		DFND	49	8,115	0	0
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	2,492	50,121	SH		DFND	15	50,121	0	0
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	66	1,332	SH		DFND	4	1,332	0	0
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	2,632	52,930	SH		DFND	49	52,930	0	0
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	1,039	20,890	SH		OTR	49	16,270	4,620	0
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	3,206	64,482	SH		DFND	59	61,345	0	3,137
ASTRAZENECA PLC-SPONS ADR	SPON	046353108	0	1	SH		DFND		1	0	0
ASTRONICS CORP	COM	046433108	498	27,580	SH		DFND	15	5,508	0	22,072
ASTRONICS CORP	COM	046433108	1,066	59,091	SH		DFND	48	59,091	0	0
ASTRONICS CORP	COM	046433108	55	3,030	SH		DFND	49	3,030	0	0
ASURE SOFTWARE INC	COM	04649U102	72	9,401	SH		DFND	15	1,540	0	7,861
ASURE SOFTWARE INC	COM	04649U102	356	46,589	SH		DFND	48	46,589	0	0
ASURE SOFTWARE INC	COM	04649U102	11	1,425	SH		DFND	49	1,425	0	0
ASURE SOFTWARE INC	COM	04649U102	0	1	SH		DFND		1	0	0
AT HOME GROUP INC	COM	04650Y100	5,900	205,578	SH		DFND	15	159,493	0	46,085
AT HOME GROUP INC	COM	04650Y100	3,893	135,658	SH		DFND	48	135,658	0	0
AT HOME GROUP INC	COM	04650Y100	162	5,655	SH		DFND	49	5,655	0	0
AT&T INC	COM	00206R102	19,363	639,688	SH		DFND	15,22	639,688	0	0
AT&T INC	COM	00206R102	18,095	597,793	SH		OTR	15,119	597,793	0	0
AT&T INC	COM	00206R102	7,295	240,997	SH		OTR	15,118	240,997	0	0
AT&T INC	COM	00206R102	940,024	31,054,644	SH		DFND	15	18,998,102	0	12,056,542
AT&T INC	COM	00206R102	67,608	2,233,500	SH		DFND	22	2,233,500	0	0
AT&T INC	COM	00206R102	2,901	95,828	SH		DFND	4	95,828	0	0
AT&T INC	COM	00206R102	1,445	47,732	SH		OTR	4	26,005	21,727	0
AT&T INC	COM	00206R102	69	2,289	SH		DFND	40	2,289	0	0
AT&T INC	COM	00206R102	636,515	21,027,913	SH		DFND	48	18,869,430	0	2,158,483
AT&T INC	COM	00206R102	107,349	3,546,378	SH		DFND	49	3,483,969	0	62,409
AT&T INC	COM	00206R102	24,126	797,018	SH		OTR	49	451,410	309,424	36,184
AT&T INC	COM	00206R102	16,755	553,506	SH		DFND	59	534,048	0	19,458
AT&T INC	COM	00206R102	0	1	SH		DFND		1	0	0
ATARA BIOTHERAPEUTICS INC	COM	046513107	1,511	105,212	SH		DFND	15	65,735	0	39,477
ATARA BIOTHERAPEUTICS INC	COM	046513107	2,724	189,708	SH		DFND	48	189,708	0	0
ATARA BIOTHERAPEUTICS INC	COM	046513107	16	1,140	SH		OTR	49	0	1,140	0
ATARA BIOTHERAPEUTICS INC	COM	046513107	0	28	SH		DFND	49	28	0	0
ATEA PHARMACEUTICALS INC	COM	04683R106	613	9,920	SH		DFND	15	3,710	0	6,210
ATEA PHARMACEUTICALS INC	COM	04683R106	2,119	34,317	SH		DFND	48	34,317	0	0
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	8,920	176,994	SH		DFND	15	118,828	0	58,166
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	525	10,416	SH		OTR	15,119	10,416	0	0
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	7	129	SH		OTR	4	0	0	129
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	11,111	220,457	SH		DFND	48	183,838	0	36,619
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	4	70	SH		OTR	49	70	0	0
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	93	1,851	SH		DFND	49	1,851	0	0
ATHENE HOLDING LTD-CLASS A	CL A	G0684D107	901	17,880	SH		DFND	59	17,880	0	0
ATHENEX INC	COM	04685N103	327	76,006	SH		DFND	15	46,774	0	29,232
ATHENEX INC	COM	04685N103	755	175,467	SH		DFND	48	175,467	0	0
ATHERSYS INC	COM	04744L106	423	234,767	SH		DFND	15	172,009	0	62,758
ATHERSYS INC	COM	04744L106	875	485,870	SH		DFND	48	485,870	0	0
ATHIRA PHARMA INC	COM	04746L104	92	5,015	SH		DFND	15	1,867	0	3,148
ATHIRA PHARMA INC	COM	04746L104	564	30,663	SH		DFND	48	30,663	0	0
ATKORE INC	COM	047649108	5,795	80,597	SH		DFND	15	64,644	0	15,953
ATKORE INC	COM	047649108	1,728	24,030	SH		OTR	22,69	24,030	0	0
ATKORE INC	COM	047649108	8,936	124,286	SH		DFND	48	124,286	0	0
ATKORE INC	COM	047649108	194	2,692	SH		DFND	49	2,692	0	0
ATKORE INC	COM	047649108	606	8,434	SH		DFND	59	8,243	0	191
ATLANTIC CAPITAL BANCSHARES	COM	048269203	583	24,180	SH		DFND	15	6,408	0	17,772
ATLANTIC CAPITAL BANCSHARES	COM	048269203	1,171	48,583	SH		DFND	48	48,583	0	0
ATLANTIC POWER CORP	COM	04878Q863	445	153,917	SH		DFND	15	52,491	0	101,426
ATLANTIC POWER CORP	COM	04878Q863	916	316,990	SH		OTR	22,83	316,990	0	0
ATLANTIC POWER CORP	COM	04878Q863	747	258,649	SH		DFND	48	258,649	0	0
ATLANTIC POWER CORP	COM	04878Q863	2	711	SH		DFND	49	711	0	0
ATLANTIC POWER CORP	COM	04878Q863	0	1	SH		DFND	59	1	0	0
ATLANTIC UNION BANKSHARES CO	COM	04911A107	11,345	295,739	SH		DFND	15	224,657	0	71,082
ATLANTIC UNION BANKSHARES CO	COM	04911A107	7,379	192,368	SH		DFND	48	192,368	0	0
ATLANTIC UNION BANKSHARES CO	COM	04911A107	21	550	SH		DFND	49	550	0	0
ATLANTIC UNION BANKSHARES CO	COM	04911A107	0	1	SH		DFND		1	0	0
ATLANTICUS HOLDINGS CORP	COM	04914Y102	220	7,257	SH		DFND	15	6,879	0	378
ATLANTICUS HOLDINGS CORP	COM	04914Y102	382	12,588	SH		DFND	48	12,588	0	0
ATLAS AIR WORLDWIDE HLDG	NOTE	049164BH8	744	700,000	SH		DFND	15	700,000	0	0
ATLAS AIR WORLDWIDE HLDG	NOTE	049164BJ4	211	175,000	SH		DFND	15	175,000	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	12,195	201,764	SH		DFND	15	179,087	0	22,677
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	137	2,262	SH		OTR	15,119	2,262	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	4,205	69,568	SH		DFND	22	69,568	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	24	395	SH		DFND	4	395	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	4,093	67,724	SH		DFND	48	67,724	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	870	14,397	SH		DFND	49	14,397	0	0
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	61,681	292,661	SH		DFND	15	220,799	0	71,862
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	4,317	20,484	SH		DFND	22	20,484	0	0
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	2,053	9,739	SH		DFND	4	9,739	0	0
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	128	605	SH		OTR	4	311	294	0
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	85,320	404,822	SH		DFND	48	364,096	0	40,726
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	61,899	293,695	SH		DFND	49	289,695	147	3,853
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	2,965	14,070	SH		OTR	49	10,826	3,230	14
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	2,022	9,592	SH		DFND	59	9,317	0	275
ATLASSIAN CORP PLC-CLASS A	CL A	G06242104	0	1	SH		DFND		1	0	0
ATMOS ENERGY CORP	COM	049560105	492	4,977	SH		OTR	15,119	4,977	0	0
ATMOS ENERGY CORP	COM	049560105	1,105	11,178	SH		DFND	15,22	11,178	0	0
ATMOS ENERGY CORP	COM	049560105	45,693	462,241	SH		DFND	15	261,877	0	200,364
ATMOS ENERGY CORP	COM	049560105	435	4,401	SH		OTR	15,118	4,401	0	0
ATMOS ENERGY CORP	COM	049560105	2,367	23,943	SH		DFND	22	23,943	0	0
ATMOS ENERGY CORP	COM	049560105	167	1,694	SH		DFND	4	1,694	0	0
ATMOS ENERGY CORP	COM	049560105	39,601	400,620	SH		DFND	48	363,027	0	37,593
ATMOS ENERGY CORP	COM	049560105	3,085	31,205	SH		DFND	49	31,205	0	0
ATMOS ENERGY CORP	COM	049560105	175	1,771	SH		OTR	49	0	180	1,591
ATMOS ENERGY CORP	COM	049560105	0	1	SH		DFND		1	0	0
ATN INTERNATIONAL INC	COM	00215F107	4,180	85,094	SH		DFND	15	74,010	0	11,084
ATN INTERNATIONAL INC	COM	00215F107	1,384	28,177	SH		DFND	22	28,177	0	0
ATN INTERNATIONAL INC	COM	00215F107	2,145	43,664	SH		OTR	22,121	43,664	0	0
ATN INTERNATIONAL INC	COM	00215F107	4	90	SH		DFND	4	90	0	0
ATN INTERNATIONAL INC	COM	00215F107	1,391	28,311	SH		DFND	48	28,311	0	0
ATN INTERNATIONAL INC	COM	00215F107	36	734	SH		DFND	49	734	0	0
ATOMERA INC	COM	04965B100	218	8,885	SH		DFND	15	2,629	0	6,256
ATOMERA INC	COM	04965B100	1,610	65,701	SH		DFND	48	65,701	0	0
ATOMERA INC	COM	04965B100	99	4,045	SH		DFND	49	4,045	0	0
ATOSSA THERAPEUTICS INC	COM	04962H506	76	35,927	SH		DFND	48	35,927	0	0
ATRECA INC - A	CL A	04965G109	593	38,709	SH		DFND	15	28,658	0	10,051
ATRECA INC - A	CL A	04965G109	1,038	67,733	SH		DFND	48	67,733	0	0
ATRICURE INC	COM	04963C209	6,261	95,559	SH		DFND	15	79,227	0	16,332
ATRICURE INC	COM	04963C209	3,228	49,263	SH		DFND	22	49,263	0	0
ATRICURE INC	COM	04963C209	7,498	114,444	SH		DFND	48	114,444	0	0
ATRICURE INC	COM	04963C209	46	700	SH		OTR	49	700	0	0
ATRICURE INC	COM	04963C209	199	3,044	SH		DFND	49	3,044	0	0
ATRION CORPORATION	COM	049904105	4,122	6,427	SH		DFND	15	5,828	0	599
ATRION CORPORATION	COM	049904105	2,180	3,400	SH		DFND	48	3,400	0	0
ATRION CORPORATION	COM	049904105	38	59	SH		DFND	49	59	0	0
ATRION CORPORATION	COM	049904105	1	1	SH		DFND		1	0	0
AUBURN NATL BANCORPORATION	COM	050473107	280	7,302	SH		DFND	15	5,927	0	1,375
AUBURN NATL BANCORPORATION	COM	050473107	177	4,609	SH		DFND	48	4,609	0	0
AUDIOCODES LTD	ORD	M15342104	453	16,799	SH		DFND	15	16,799	0	0
AURINIA PHARMACEUTICALS INC	COM	05156V102	501	38,600	SH		DFND	15	38,600	0	0
AURINIA PHARMACEUTICALS INC	COM	05156V102	43	3,281	SH		DFND	48	3,281	0	0
AURINIA PHARMACEUTICALS INC	COM	05156V102	134	10,319	SH		DFND	49	10,319	0	0
AUTODESK INC	COM	052769106	176	635	SH		OTR	15,119	635	0	0
AUTODESK INC	COM	052769106	260,316	939,262	SH		DFND	15	566,875	0	372,387
AUTODESK INC	COM	052769106	5,446	19,651	SH		DFND	15,22	19,651	0	0
AUTODESK INC	COM	052769106	2,057	7,421	SH		OTR	15,118	7,421	0	0
AUTODESK INC	COM	052769106	10,645	38,407	SH		DFND	22	38,407	0	0
AUTODESK INC	COM	052769106	363	1,308	SH		DFND	4	1,308	0	0
AUTODESK INC	COM	052769106	1	4	SH		OTR	4	0	4	0
AUTODESK INC	COM	052769106	183,804	663,195	SH		DFND	48	595,678	0	67,517
AUTODESK INC	COM	052769106	17,811	64,265	SH		DFND	49	63,975	0	290
AUTODESK INC	COM	052769106	504	1,817	SH		OTR	49	363	1,096	358
AUTODESK INC	COM	052769106	1,012	3,653	SH		DFND	59	3,492	0	161
AUTODESK INC	COM	052769106	0	1	SH		DFND		1	0	0
AUTOHOME INC-ADR	SP A	05278C107	17,558	188,254	SH		DFND	15	186,957	0	1,297
AUTOHOME INC-ADR	SP A	05278C107	7,078	75,882	SH		OTR	40,48	75,882	0	0
AUTOHOME INC-ADR	SP A	05278C107	13,287	142,454	SH		DFND	40,22	142,454	0	0
AUTOHOME INC-ADR	SP A	05278C107	32,370	347,055	SH		DFND	40	333,359	0	13,696
AUTOHOME INC-ADR	SP A	05278C107	4,974	53,332	SH		DFND	48	53,332	0	0
AUTOHOME INC-ADR	SP A	05278C107	1	14	SH		OTR	49	0	14	0
AUTOHOME INC-ADR	SP A	05278C107	1,615	17,313	SH		DFND	59	17,049	0	264
AUTOHOME INC-ADR	SP A	05278C107	0	1	SH		DFND		1	0	0
AUTOLIV INC	COM	052800109	397	4,281	SH		OTR	15,119	4,281	0	0
AUTOLIV INC	COM	052800109	11,293	121,689	SH		DFND	15	118,844	0	2,845
AUTOLIV INC	COM	052800109	5,149	55,488	SH		DFND	48	55,488	0	0
AUTOLIV INC	COM	052800109	279	3,003	SH		DFND	49	2,978	0	25
AUTOLIV INC	COM	052800109	802	8,645	SH		OTR	49	21	335	8,289
AUTOLIV INC	COM	052800109	0	1	SH		DFND		1	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	549,030	2,913,087	SH		DFND	15	2,187,751	0	725,336
AUTOMATIC DATA PROCESSING	COM	053015103	2,723	14,450	SH		OTR	15,118	14,450	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	1,671	8,867	SH		OTR	15,119	8,867	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	7,227	38,347	SH		DFND	15,22	38,347	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	14,196	75,321	SH		DFND	22	75,321	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	509	2,700	SH		OTR	4	1,900	800	0
AUTOMATIC DATA PROCESSING	COM	053015103	3,415	18,118	SH		DFND	4	18,118	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	46	245	SH		DFND	40	245	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	241,727	1,282,577	SH		DFND	48	1,149,622	0	132,955
AUTOMATIC DATA PROCESSING	COM	053015103	22,564	119,722	SH		OTR	49	62,716	57,006	0
AUTOMATIC DATA PROCESSING	COM	053015103	60,612	321,602	SH		DFND	49	309,869	0	11,733
AUTOMATIC DATA PROCESSING	COM	053015103	21,103	111,968	SH		OTR	51,48	111,968	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	170,355	903,883	SH		OTR	51,94	903,883	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	3,242	17,200	SH		OTR	51	17,200	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	43,989	233,400	SH		DFND	51,22	233,400	0	0
AUTOMATIC DATA PROCESSING	COM	053015103	1,333,698	7,076,448	SH		DFND	51	5,021,808	0	2,054,640
AUTOMATIC DATA PROCESSING	COM	053015103	26,552	140,882	SH		DFND	59	133,909	0	6,973
AUTOMATIC DATA PROCESSING	COM	053015103	234	1,240	SH		DFND		1,240	0	0
AUTONATION INC	COM	05329W102	677	7,267	SH		OTR	15,119	7,267	0	0
AUTONATION INC	COM	05329W102	27,995	300,307	SH		DFND	15	273,607	0	26,700
AUTONATION INC	COM	05329W102	345	3,700	SH		DFND	15,22	3,700	0	0
AUTONATION INC	COM	05329W102	6,015	64,529	SH		DFND	22	64,529	0	0
AUTONATION INC	COM	05329W102	54	580	SH		DFND	4	580	0	0
AUTONATION INC	COM	05329W102	20,474	219,634	SH		DFND	48	202,362	0	17,272
AUTONATION INC	COM	05329W102	55	590	SH		OTR	49	0	590	0
AUTONATION INC	COM	05329W102	2,902	31,136	SH		DFND	49	31,016	0	120
AUTONATION INC	COM	05329W102	311	3,338	SH		DFND	59	3,338	0	0
AUTOWEB INC	COM	05335B100	116	43,298	SH		DFND	48	43,298	0	0
AUTOZONE INC	COM	053332102	1,050	748	SH		OTR	15,118	748	0	0
AUTOZONE INC	COM	053332102	2,797	1,992	SH		DFND	15,22	1,992	0	0
AUTOZONE INC	COM	053332102	862	614	SH		OTR	15,119	614	0	0
AUTOZONE INC	COM	053332102	120,738	85,977	SH		DFND	15	49,942	0	36,035
AUTOZONE INC	COM	053332102	6,378	4,542	SH		OTR	22,80	4,542	0	0
AUTOZONE INC	COM	053332102	12,658	9,014	SH		DFND	22	9,014	0	0
AUTOZONE INC	COM	053332102	268	191	SH		DFND	4	191	0	0
AUTOZONE INC	COM	053332102	10	7	SH		OTR	4	0	0	7
AUTOZONE INC	COM	053332102	93,594	66,648	SH		DFND	48	59,647	0	7,001
AUTOZONE INC	COM	053332102	9,799	6,978	SH		DFND	49	6,933	0	45
AUTOZONE INC	COM	053332102	125	89	SH		OTR	49	53	36	0
AUTOZONE INC	COM	053332102	2,603	1,854	SH		DFND	59	1,787	0	67
AUTOZONE INC	COM	053332102	1	1	SH		DFND		1	0	0
AVALARA INC	COM	05338G106	36,372	272,589	SH		DFND	15	230,274	0	42,315
AVALARA INC	COM	05338G106	19,907	149,193	SH		DFND	48	123,806	0	25,387
AVALARA INC	COM	05338G106	5	40	SH		OTR	49	0	40	0
AVALARA INC	COM	05338G106	3,935	29,488	SH		DFND	49	29,488	0	0
AVALARA INC	COM	05338G106	343	2,571	SH		DFND	59	2,482	0	89
AVALON GLOBOCARE CORP	COM	05344R104	2	1,456	SH		DFND	15	0	0	1,456
AVALON GLOBOCARE CORP	COM	05344R104	30	27,671	SH		DFND	48	27,671	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	2,309	12,516	SH		DFND	15,22	12,516	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	882	4,780	SH		OTR	15,119	4,780	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	94,356	511,389	SH		DFND	15	285,118	0	226,271
AVALONBAY COMMUNITIES INC	COM	053484101	867	4,699	SH		OTR	15,118	4,699	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	5,778	31,314	SH		DFND	22	31,314	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	183	993	SH		DFND	4	993	0	0
AVALONBAY COMMUNITIES INC	COM	053484101	98,491	533,796	SH		DFND	48	491,321	0	42,475
AVALONBAY COMMUNITIES INC	COM	053484101	359	1,944	SH		OTR	49	536	1,408	0
AVALONBAY COMMUNITIES INC	COM	053484101	5,324	28,856	SH		DFND	49	28,788	0	68
AVALONBAY COMMUNITIES INC	COM	053484101	42	228	SH		DFND	59	228	0	0
AVANGRID INC	COM	05351W103	18,025	361,871	SH		DFND	15	328,496	0	33,375
AVANGRID INC	COM	05351W103	184	3,691	SH		OTR	15,119	3,691	0	0
AVANGRID INC	COM	05351W103	4,931	99,005	SH		DFND	48	82,487	0	16,518
AVANGRID INC	COM	05351W103	225	4,510	SH		DFND	49	4,510	0	0
AVANGRID INC	COM	05351W103	2	41	SH		OTR	49	0	41	0
AVANGRID INC	COM	05351W103	25	493	SH		DFND		493	0	0
AVANOS MEDICAL INC	COM	05350V106	120	2,740	SH		DFND	15,22	2,740	0	0
AVANOS MEDICAL INC	COM	05350V106	16	376	SH		OTR	15,119	376	0	0
AVANOS MEDICAL INC	COM	05350V106	9,653	220,701	SH		DFND	15	178,583	0	42,118
AVANOS MEDICAL INC	COM	05350V106	1,922	43,950	SH		OTR	22,121	43,950	0	0
AVANOS MEDICAL INC	COM	05350V106	2,431	55,579	SH		DFND	22	55,579	0	0
AVANOS MEDICAL INC	COM	05350V106	3	70	SH		DFND	4	70	0	0
AVANOS MEDICAL INC	COM	05350V106	9,254	211,562	SH		DFND	48	211,562	0	0
AVANOS MEDICAL INC	COM	05350V106	894	20,448	SH		DFND	49	20,328	0	120
AVANOS MEDICAL INC	COM	05350V106	59	1,353	SH		OTR	49	885	468	0
AVANOS MEDICAL INC	COM	05350V106	0	1	SH		DFND		1	0	0
AVANTOR INC	COM	05352A100	37,071	1,281,414	SH		DFND	15	1,029,149	0	252,265
AVANTOR INC	COM	05352A100	163	5,622	SH		OTR	15,119	5,622	0	0
AVANTOR INC	COM	05352A100	4,261	147,272	SH		OTR	22,80	147,272	0	0
AVANTOR INC	COM	05352A100	849	29,353	SH		OTR	22,78	29,353	0	0
AVANTOR INC	COM	05352A100	2,084	72,030	SH		DFND	22	72,030	0	0
AVANTOR INC	COM	05352A100	8	262	SH		OTR	4	0	0	262
AVANTOR INC	COM	05352A100	27,965	966,655	SH		DFND	48	812,515	0	154,140
AVANTOR INC	COM	05352A100	141	4,860	SH		DFND	49	4,860	0	0
AVANTOR INC	COM	05352A100	1,059	36,594	SH		OTR	49	24,220	12,374	0
AVANTOR INC	COM	05352A100	1,332	46,055	SH		DFND	59	43,542	0	2,513
AVAYA HOLDINGS CORP	COM	05351X101	151	5,373	SH		OTR	15,119	5,373	0	0
AVAYA HOLDINGS CORP	COM	05351X101	2,615	93,304	SH		DFND	15	63,817	0	29,487
AVAYA HOLDINGS CORP	COM	05351X101	1,295	46,202	SH		OTR	22,69	46,202	0	0
AVAYA HOLDINGS CORP	COM	05351X101	3,544	126,450	SH		OTR	22,96	126,450	0	0
AVAYA HOLDINGS CORP	COM	05351X101	6,051	215,892	SH		DFND	48	215,892	0	0
AVAYA HOLDINGS CORP	COM	05351X101	0	10	SH		DFND		10	0	0
AVENUE THERAPEUTICS INC	COM	05360L205	8	1,262	SH		DFND	15	764	0	498
AVENUE THERAPEUTICS INC	COM	05360L205	79	12,936	SH		DFND	48	12,936	0	0
AVENUE THERAPEUTICS INC	COM	05360L205	4	650	SH		DFND	49	650	0	0
AVEO PHARMACEUTICALS INC	COM	053588307	55	7,451	SH		DFND	15	1,559	0	5,892
AVEO PHARMACEUTICALS INC	COM	053588307	351	47,943	SH		DFND	48	47,943	0	0
AVEO PHARMACEUTICALS INC	COM	053588307	27	3,655	SH		DFND	49	3,655	0	0
AVERY DENNISON CORP	COM	053611109	95,252	518,658	SH		DFND	15	382,548	0	136,110
AVERY DENNISON CORP	COM	053611109	512	2,789	SH		OTR	15,118	2,789	0	0
AVERY DENNISON CORP	COM	053611109	408	2,224	SH		OTR	15,119	2,224	0	0
AVERY DENNISON CORP	COM	053611109	1,367	7,441	SH		DFND	15,22	7,441	0	0
AVERY DENNISON CORP	COM	053611109	2,024	11,023	SH		OTR	22,80	11,023	0	0
AVERY DENNISON CORP	COM	053611109	5,631	30,660	SH		DFND	22	30,660	0	0
AVERY DENNISON CORP	COM	053611109	12	65	SH		DFND	4	65	0	0
AVERY DENNISON CORP	COM	053611109	4	23	SH		OTR	4	0	0	23
AVERY DENNISON CORP	COM	053611109	48,982	266,716	SH		DFND	48	241,195	0	25,521
AVERY DENNISON CORP	COM	053611109	5,742	31,267	SH		DFND	49	31,267	0	0
AVERY DENNISON CORP	COM	053611109	987	5,377	SH		OTR	49	3,384	1,147	846
AVERY DENNISON CORP	COM	053611109	1,019	5,548	SH		DFND	59	5,303	0	245
AVERY DENNISON CORP	COM	053611109	0	1	SH		DFND		1	0	0
AVIAT NETWORKS INC	COM	05366Y201	928	13,061	SH		DFND	15	13,061	0	0
AVIAT NETWORKS INC	COM	05366Y201	179	2,514	SH		DFND	48	2,514	0	0
AVID BIOSERVICES INC	COM	05368M106	1,069	58,619	SH		DFND	15	37,954	0	20,665
AVID BIOSERVICES INC	COM	05368M106	2,434	133,505	SH		DFND	48	133,505	0	0
AVID BIOSERVICES INC	COM	05368M106	0	6	SH		DFND		6	0	0
AVID TECHNOLOGY INC	COM	05367P100	1,070	50,709	SH		DFND	15	34,761	0	15,948
AVID TECHNOLOGY INC	COM	05367P100	1,706	80,810	SH		DFND	48	80,810	0	0
AVID TECHNOLOGY INC	COM	05367P100	84	4,000	SH		DFND	49	4,000	0	0
AVIDITY BIOSCIENCES INC	COM	05370A108	562	25,770	SH		DFND	15	17,378	0	8,392
AVIDITY BIOSCIENCES INC	COM	05370A108	1,280	58,681	SH		DFND	48	58,681	0	0
AVIENT CORP	COM	05368V106	194	4,099	SH		OTR	15,119	4,099	0	0
AVIENT CORP	COM	05368V106	27,620	584,302	SH		DFND	15	503,618	0	80,684
AVIENT CORP	COM	05368V106	4,589	97,081	SH		OTR	22,96	97,081	0	0
AVIENT CORP	COM	05368V106	4,580	96,884	SH		DFND	22	96,884	0	0
AVIENT CORP	COM	05368V106	69	1,465	SH		DFND	4	1,465	0	0
AVIENT CORP	COM	05368V106	19,357	409,490	SH		DFND	48	409,490	0	0
AVIENT CORP	COM	05368V106	8	174	SH		OTR	49	0	174	0
AVIENT CORP	COM	05368V106	2,254	47,690	SH		DFND	49	47,500	0	190
AVIENT CORP	COM	05368V106	0	1	SH		DFND		1	0	0
AVINGER INC	COM	053734604	69	45,904	SH		DFND	48	45,904	0	0
AVIS BUDGET GROUP INC	COM	053774105	18,759	258,601	SH		DFND	15	239,435	0	19,166
AVIS BUDGET GROUP INC	COM	053774105	783	10,794	SH		OTR	15,119	10,794	0	0
AVIS BUDGET GROUP INC	COM	053774105	268	3,690	SH		DFND	15,22	3,690	0	0
AVIS BUDGET GROUP INC	COM	053774105	4,422	60,960	SH		DFND	22	60,960	0	0
AVIS BUDGET GROUP INC	COM	053774105	1,289	17,769	SH		OTR	22,69	17,769	0	0
AVIS BUDGET GROUP INC	COM	053774105	2	30	SH		DFND	4	30	0	0
AVIS BUDGET GROUP INC	COM	053774105	17,703	244,039	SH		DFND	48	244,039	0	0
AVIS BUDGET GROUP INC	COM	053774105	1,363	18,793	SH		DFND	49	18,703	0	90
AVIS BUDGET GROUP INC	COM	053774105	6	78	SH		OTR	49	7	71	0
AVIS BUDGET GROUP INC	COM	053774105	0	1	SH		DFND		1	0	0
AVISTA CORP	COM	05379B107	7,594	159,028	SH		OTR	15,66	159,028	0	0
AVISTA CORP	COM	05379B107	215	4,496	SH		OTR	15,119	4,496	0	0
AVISTA CORP	COM	05379B107	50,557	1,058,777	SH		DFND	15	948,841	0	109,936
AVISTA CORP	COM	05379B107	12,511	262,009	SH		DFND	22	262,009	0	0
AVISTA CORP	COM	05379B107	175	3,670	SH		OTR	22,87	3,670	0	0
AVISTA CORP	COM	05379B107	30	630	SH		DFND	4	630	0	0
AVISTA CORP	COM	05379B107	9,365	196,117	SH		DFND	48	196,117	0	0
AVISTA CORP	COM	05379B107	2,365	49,527	SH		DFND	49	49,527	0	0
AVISTA CORP	COM	05379B107	210	4,395	SH		OTR	49	4,395	0	0
AVISTA CORP	COM	05379B107	0	1	SH		DFND		1	0	0
AVITA MEDICAL INC	COM	05380C102	234	11,806	SH		DFND	48	11,806	0	0
AVNET INC	COM	053807103	508	12,230	SH		DFND	15,22	12,230	0	0
AVNET INC	COM	053807103	530	12,759	SH		OTR	15,119	12,759	0	0
AVNET INC	COM	053807103	28,230	680,077	SH		DFND	15	624,168	0	55,909
AVNET INC	COM	053807103	6,959	167,652	SH		DFND	22	167,652	0	0
AVNET INC	COM	053807103	52	1,252	SH		DFND	4	1,252	0	0
AVNET INC	COM	053807103	15,995	385,318	SH		DFND	48	354,615	0	30,703
AVNET INC	COM	053807103	2,001	48,211	SH		DFND	49	48,011	0	200
AVNET INC	COM	053807103	8	183	SH		OTR	49	56	127	0
AVROBIO INC	COM	05455M100	6,697	527,751	SH		DFND	15,22	527,751	0	0
AVROBIO INC	COM	05455M100	1,576	124,216	SH		DFND	15	100,389	0	23,827
AVROBIO INC	COM	05455M100	833	65,625	SH		DFND	22	65,625	0	0
AVROBIO INC	COM	05455M100	195	15,376	SH		DFND	42	15,376	0	0
AVROBIO INC	COM	05455M100	1,553	122,380	SH		DFND	48	122,380	0	0
AWARE INC/MASS	COM	05453N100	61	16,773	SH		DFND	48	16,773	0	0
AWARE INC/MASS	COM	05453N100	0	1	SH		DFND		1	0	0
AXALTA COATING SYSTEMS LTD	COM	G0750C108	245	8,270	SH		OTR	15,119	8,270	0	0
AXALTA COATING SYSTEMS LTD	COM	G0750C108	9,803	331,422	SH		DFND	15	218,187	0	113,235
AXALTA COATING SYSTEMS LTD	COM	G0750C108	35	1,191	SH		OTR	4	0	0	1,191
AXALTA COATING SYSTEMS LTD	COM	G0750C108	0	15	SH		DFND	4	15	0	0
AXALTA COATING SYSTEMS LTD	COM	G0750C108	11,237	379,879	SH		DFND	48	318,472	0	61,407
AXALTA COATING SYSTEMS LTD	COM	G0750C108	2,062	69,696	SH		OTR	49	69,696	0	0
AXALTA COATING SYSTEMS LTD	COM	G0750C108	33	1,122	SH		DFND	49	1,122	0	0
AXALTA COATING SYSTEMS LTD	COM	G0750C108	980	33,124	SH		DFND	59	31,659	0	1,465
AXALTA COATING SYSTEMS LTD	COM	G0750C108	0	1	SH		DFND		1	0	0
AXCELIS TECHNOLOGIES INC	COM	054540208	6,837	166,394	SH		DFND	15	154,204	0	12,190
AXCELIS TECHNOLOGIES INC	COM	054540208	3,549	86,367	SH		DFND	22	86,367	0	0
AXCELIS TECHNOLOGIES INC	COM	054540208	794	19,330	SH		OTR	22,83	19,330	0	0
AXCELIS TECHNOLOGIES INC	COM	054540208	3,602	87,658	SH		DFND	48	87,658	0	0
AXCELIS TECHNOLOGIES INC	COM	054540208	89	2,175	SH		DFND	49	2,175	0	0
AXCELLA HEALTH INC	COM	05454B105	200	41,912	SH		DFND	15	40,817	0	1,095
AXCELLA HEALTH INC	COM	05454B105	209	43,856	SH		DFND	48	43,856	0	0
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	4,946	99,774	SH		DFND	15	57,638	0	42,136
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	231	4,656	SH		OTR	15,119	4,656	0	0
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	1,090	21,980	SH		OTR	22,83	21,980	0	0
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	7,433	149,943	SH		DFND	48	123,860	0	26,083
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	301	6,076	SH		OTR	49	6,013	63	0
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	545	11,003	SH		DFND	49	11,003	0	0
AXIS CAPITAL HOLDINGS LTD	SHS	G0692U109	0	1	SH		DFND		1	0	0
AXOGEN INC	COM	05463X106	968	47,768	SH		DFND	15	33,357	0	14,411
AXOGEN INC	COM	05463X106	1,733	85,540	SH		DFND	48	85,540	0	0
AXON ENTERPRISE INC	COM	05464C101	72	504	SH		OTR	15,119	504	0	0
AXON ENTERPRISE INC	COM	05464C101	219	1,540	SH		DFND	15,22	1,540	0	0
AXON ENTERPRISE INC	COM	05464C101	37,854	265,791	SH		DFND	15	234,423	0	31,368
AXON ENTERPRISE INC	COM	05464C101	8,207	57,624	SH		OTR	22,71	57,624	0	0
AXON ENTERPRISE INC	COM	05464C101	10,351	72,681	SH		DFND	22	72,681	0	0
AXON ENTERPRISE INC	COM	05464C101	125	876	SH		DFND	4	876	0	0
AXON ENTERPRISE INC	COM	05464C101	35,653	250,339	SH		DFND	48	231,065	0	19,274
AXON ENTERPRISE INC	COM	05464C101	113	791	SH		OTR	49	0	41	750
AXON ENTERPRISE INC	COM	05464C101	3,933	27,617	SH		DFND	49	27,498	0	119
AXONICS MODULATION TECHNOLOG	COM	05465P101	1,915	31,971	SH		DFND	15	22,385	0	9,586
AXONICS MODULATION TECHNOLOG	COM	05465P101	4,522	75,499	SH		DFND	48	75,499	0	0
AXONICS MODULATION TECHNOLOG	COM	05465P101	19	310	SH		DFND	49	310	0	0
AXOS FINANCIAL INC	COM	05465C100	14,421	306,755	SH		DFND	15	259,978	0	46,777
AXOS FINANCIAL INC	COM	05465C100	6,242	132,770	SH		DFND	22	132,770	0	0
AXOS FINANCIAL INC	COM	05465C100	2,500	53,174	SH		OTR	22,87	53,174	0	0
AXOS FINANCIAL INC	COM	05465C100	1,609	34,236	SH		OTR	22,71	34,236	0	0
AXOS FINANCIAL INC	COM	05465C100	41	879	SH		DFND	4	879	0	0
AXOS FINANCIAL INC	COM	05465C100	6,931	147,443	SH		DFND	48	147,443	0	0
AXOS FINANCIAL INC	COM	05465C100	1,225	26,061	SH		DFND	49	26,061	0	0
AXSOME THERAPEUTICS INC	COM	05464T104	2,181	38,519	SH		DFND	15	28,410	0	10,109
AXSOME THERAPEUTICS INC	COM	05464T104	313	5,521	SH		OTR	22,78	5,521	0	0
AXSOME THERAPEUTICS INC	COM	05464T104	396	6,990	SH		OTR	22,104	6,990	0	0
AXSOME THERAPEUTICS INC	COM	05464T104	4,565	80,626	SH		DFND	48	80,626	0	0
AXSOME THERAPEUTICS INC	COM	05464T104	113	1,994	SH		DFND	49	1,994	0	0
AXSOME THERAPEUTICS INC	COM	05464T104	641	11,324	SH		OTR	49	0	11,324	0
AXT INC	COM	00246W103	505	43,332	SH		DFND	15	11,372	0	31,960
AXT INC	COM	00246W103	1,060	90,922	SH		DFND	48	90,922	0	0
AYRO INC	COM	054748108	135	20,852	SH		DFND	48	20,852	0	0
AYRO INC	COM	054748108	0	52	SH		DFND		52	0	0
AYTU BIOPHARMA INC	COM	054754874	59	7,711	SH		DFND	15	0	0	7,711
AYTU BIOPHARMA INC	COM	054754874	296	38,936	SH		DFND	48	38,936	0	0
AYTU BIOPHARMA INC	COM	054754874	1	185	SH		DFND	49	185	0	0
AYTU BIOPHARMA INC	COM	054754874	0	1	SH		DFND		1	0	0
AZEK CO INC/THE	CL A	05478C105	28,994	689,504	SH		DFND	15,22	689,504	0	0
AZEK CO INC/THE	CL A	05478C105	6,532	155,345	SH		DFND	15	95,918	0	59,427
AZEK CO INC/THE	CL A	05478C105	1,854	44,096	SH		DFND	22	44,096	0	0
AZEK CO INC/THE	CL A	05478C105	705	16,771	SH		DFND	42	16,771	0	0
AZEK CO INC/THE	CL A	05478C105	8,777	208,730	SH		DFND	48	181,932	0	26,798
AZEK CO INC/THE	CL A	05478C105	57	1,353	SH		DFND	49	1,353	0	0
AZEK CO INC/THE	CL A	05478C105	161	3,821	SH		OTR	49	3,225	596	0
AZURRX BIOPHARMA INC	COM	05502L105	874	652,026	SH		DFND	49	652,026	0	0
AZZ INC	COM	002474104	10,010	198,810	SH		DFND	15	176,530	0	22,280
AZZ INC	COM	002474104	1,069	21,230	SH		OTR	22,83	21,230	0	0
AZZ INC	COM	002474104	3,303	65,598	SH		DFND	22	65,598	0	0
AZZ INC	COM	002474104	3,430	68,117	SH		DFND	48	68,117	0	0
AZZ INC	COM	002474104	153	3,035	SH		DFND	49	3,035	0	0
B&G FOODS INC	COM	05508R106	45,520	1,465,561	SH		DFND	15	1,439,528	0	26,033
B&G FOODS INC	COM	05508R106	158	5,101	SH		OTR	15,119	5,101	0	0
B&G FOODS INC	COM	05508R106	5,167	166,353	SH		DFND	22	166,353	0	0
B&G FOODS INC	COM	05508R106	36	1,148	SH		DFND	4	1,148	0	0
B&G FOODS INC	COM	05508R106	5,222	168,141	SH		DFND	48	168,141	0	0
B&G FOODS INC	COM	05508R106	203	6,532	SH		OTR	49	6,508	24	0
B&G FOODS INC	COM	05508R106	1,936	62,315	SH		DFND	49	62,315	0	0
B&G FOODS INC	COM	05508R106	0	1	SH		DFND	59	1	0	0
B&G FOODS INC	COM	05508R106	0	1	SH		DFND		1	0	0
B. RILEY FINANCIAL INC	COM	05580M108	9,260	164,248	SH		DFND	15	146,587	0	17,661
B. RILEY FINANCIAL INC	COM	05580M108	3,319	58,875	SH		OTR	22,121	58,875	0	0
B. RILEY FINANCIAL INC	COM	05580M108	2,589	45,926	SH		DFND	48	45,926	0	0
B. RILEY FINANCIAL INC	COM	05580M108	108	1,920	SH		DFND	49	1,920	0	0
B2GOLD CORP	COM	11777Q209	40	9,200	SH		OTR	15,119	9,200	0	0
B2GOLD CORP	COM	11777Q209	5,553	1,288,441	SH		DFND	15	1,097,148	0	191,293
B2GOLD CORP	COM	11777Q209	2,314	536,900	SH		DFND	48	536,900	0	0
B2GOLD CORP	COM	11777Q209	6	1,487	SH		DFND	49	1,487	0	0
BABCOCK & WILCOX ENTERPR	COM	05614L209	179	18,904	SH		DFND	48	18,904	0	0
BADGER METER INC	COM	056525108	17,455	187,542	SH		DFND	15	177,681	0	9,861
BADGER METER INC	COM	056525108	6,977	74,967	SH		DFND	22	74,967	0	0
BADGER METER INC	COM	056525108	34	370	SH		DFND	4	370	0	0
BADGER METER INC	COM	056525108	7,161	76,939	SH		DFND	48	76,939	0	0
BADGER METER INC	COM	056525108	5,352	57,500	SH		OTR	49	0	0	57,500
BADGER METER INC	COM	056525108	612	6,574	SH		DFND	49	6,574	0	0
BADGER METER INC	COM	056525108	5	55	SH		DFND	59	55	0	0
BADGER METER INC	COM	056525108	0	1	SH		DFND		1	0	0
BAIDU INC - SPON ADR	SPON	056752108	608	2,797	SH		DFND	15,22	2,797	0	0
BAIDU INC - SPON ADR	SPON	056752108	124,372	571,694	SH		DFND	15	566,049	0	5,645
BAIDU INC - SPON ADR	SPON	056752108	52,535	241,484	SH		DFND	48	241,484	0	0
BAIDU INC - SPON ADR	SPON	056752108	329	1,513	SH		DFND	49	1,113	0	400
BAIDU INC - SPON ADR	SPON	056752108	293	1,347	SH		OTR	49	1,178	169	0
BAIDU INC - SPON ADR	SPON	056752108	5	25	SH		DFND	59	25	0	0
BAIDU INC - SPON ADR	SPON	056752108	0	1	SH		DFND		1	0	0
BAKER HUGHES CO	CL A	05722G100	78,948	3,653,320	SH		DFND	15	2,571,770	0	1,081,550
BAKER HUGHES CO	CL A	05722G100	1,374	63,571	SH		OTR	15,119	63,571	0	0
BAKER HUGHES CO	CL A	05722G100	540	24,972	SH		OTR	15,118	24,972	0	0
BAKER HUGHES CO	CL A	05722G100	1,355	62,685	SH		DFND	15,22	62,685	0	0
BAKER HUGHES CO	CL A	05722G100	10,212	472,572	SH		DFND	22	472,572	0	0
BAKER HUGHES CO	CL A	05722G100	143	6,610	SH		DFND	4	6,610	0	0
BAKER HUGHES CO	CL A	05722G100	43,142	1,996,398	SH		DFND	48	1,798,457	0	197,941
BAKER HUGHES CO	CL A	05722G100	3,950	182,779	SH		DFND	49	182,154	0	625
BAKER HUGHES CO	CL A	05722G100	465	21,536	SH		OTR	49	790	20,746	0
BAKER HUGHES CO	CL A	05722G100	688	31,849	SH		DFND	59	29,887	0	1,962
BALCHEM CORP	COM	057665200	19,105	152,344	SH		DFND	15	139,837	0	12,507
BALCHEM CORP	COM	057665200	10,464	83,436	SH		DFND	22	83,436	0	0
BALCHEM CORP	COM	057665200	1,048	8,355	SH		OTR	22,104	8,355	0	0
BALCHEM CORP	COM	057665200	2,972	23,700	SH		OTR	22,71	23,700	0	0
BALCHEM CORP	COM	057665200	39	312	SH		DFND	4	312	0	0
BALCHEM CORP	COM	057665200	10,258	81,793	SH		DFND	48	81,793	0	0
BALCHEM CORP	COM	057665200	97	775	SH		OTR	49	0	0	775
BALCHEM CORP	COM	057665200	1,557	12,418	SH		DFND	49	12,418	0	0
BALL CORP	COM	058498106	106,041	1,251,372	SH		DFND	15	726,791	0	524,581
BALL CORP	COM	058498106	537	6,334	SH		OTR	15,119	6,334	0	0
BALL CORP	COM	058498106	2,522	29,760	SH		DFND	15,22	29,760	0	0
BALL CORP	COM	058498106	936	11,046	SH		OTR	15,118	11,046	0	0
BALL CORP	COM	058498106	6,801	80,258	SH		DFND	22	80,258	0	0
BALL CORP	COM	058498106	78	921	SH		DFND	4	921	0	0
BALL CORP	COM	058498106	82,905	978,344	SH		DFND	48	880,130	0	98,214
BALL CORP	COM	058498106	3,087	36,433	SH		OTR	49	23,930	12,503	0
BALL CORP	COM	058498106	7,157	84,460	SH		DFND	49	84,275	0	185
BALL CORP	COM	058498106	137	1,615	SH		DFND	59	1,491	0	124
BALL CORP	COM	058498106	0	1	SH		DFND		1	0	0
BALLANTYNE STRONG INC	COM	058516105	39	15,129	SH		DFND	48	15,129	0	0
BALLARD POWER SYSTEMS INC	COM	058586108	7,930	325,799	SH		DFND	15	318,799	0	7,000
BALLARD POWER SYSTEMS INC	COM	058586108	49	2,000	SH		OTR	15,119	2,000	0	0
BALLARD POWER SYSTEMS INC	COM	058586108	2,821	115,900	SH		DFND	48	115,900	0	0
BALLARD POWER SYSTEMS INC	COM	058586108	24	1,000	SH		DFND	49	1,000	0	0
BALLARD POWER SYSTEMS INC	COM	058586108	0	1	SH		DFND		1	0	0
BALLY'S CORP	COM	05875B106	6,549	100,778	SH		DFND	15	67,645	0	33,133
BALLY'S CORP	COM	05875B106	5,691	87,576	SH		OTR	15,66	87,576	0	0
BALLY'S CORP	COM	05875B106	1,728	26,598	SH		OTR	22,69	26,598	0	0
BALLY'S CORP	COM	05875B106	1,603	24,664	SH		DFND	22	24,664	0	0
BALLY'S CORP	COM	05875B106	2,869	44,148	SH		DFND	48	44,148	0	0
BALLY'S CORP	COM	05875B106	0	1	SH		DFND		1	0	0
BANC OF CALIFORNIA INC	COM	05990K106	3,757	207,814	SH		DFND	15	166,988	0	40,826
BANC OF CALIFORNIA INC	COM	05990K106	2,099	116,104	SH		DFND	22	116,104	0	0
BANC OF CALIFORNIA INC	COM	05990K106	2,159	119,408	SH		DFND	48	119,408	0	0
BANC OF CALIFORNIA INC	COM	05990K106	63	3,460	SH		DFND	49	3,460	0	0
BANC OF CALIFORNIA INC	COM	05990K106	0	8	SH		DFND		8	0	0
BANCFIRST CORP	COM	05945F103	11,797	166,885	SH		DFND	15	149,560	0	17,325
BANCFIRST CORP	COM	05945F103	3,424	48,433	SH		DFND	22	48,433	0	0
BANCFIRST CORP	COM	05945F103	3,429	48,514	SH		DFND	48	48,514	0	0
BANCFIRST CORP	COM	05945F103	49	687	SH		DFND	49	687	0	0
BANCFIRST CORP	COM	05945F103	0	1	SH		DFND		1	0	0
BANCO BILBAO VIZCAYA-SP ADR	SPON	05946K101	27	5,185	SH		DFND	4	5,185	0	0
BANCO BILBAO VIZCAYA-SP ADR	SPON	05946K101	30	5,803	SH		OTR	4	0	0	5,803
BANCO BILBAO VIZCAYA-SP ADR	SPON	05946K101	624	119,579	SH		DFND	49	119,579	0	0
BANCO BILBAO VIZCAYA-SP ADR	SPON	05946K101	200	38,406	SH		OTR	49	0	2,052	36,354
BANCO BILBAO VIZCAYA-SP ADR	SPON	05946K101	1,430	273,984	SH		DFND	59	265,910	0	8,074
BANCO BRADESCO-ADR	SP A	059460303	29	6,102	SH		OTR	49	0	6,102	0
BANCO BRADESCO-ADR	SP A	059460303	1,650	351,010	SH		DFND	59	348,603	0	2,407
BANCO BRADESCO-ADR	SP A	059460303	0	1	SH		DFND		1	0	0
BANCO SANTANDER SA-SPON ADR	ADR	05964H105	26	7,459	SH		DFND	4	7,459	0	0
BANCO SANTANDER SA-SPON ADR	ADR	05964H105	919	268,037	SH		DFND	49	268,037	0	0
BANCO SANTANDER SA-SPON ADR	ADR	05964H105	331	96,379	SH		OTR	49	94,956	1,423	0
BANCO SANTANDER SA-SPON ADR	ADR	05964H105	553	161,132	SH		DFND	59	152,289	0	8,843
BANCO SANTANDER SA-SPON ADR	ADR	05964H105	0	17	SH		DFND		17	0	0
BANCORP INC/THE	COM	05969A105	2,532	122,210	SH		DFND	15	79,358	0	42,852
BANCORP INC/THE	COM	05969A105	2,880	139,020	SH		DFND	48	139,020	0	0
BANCORP INC/THE	COM	05969A105	1	63	SH		DFND	49	63	0	0
BANCORP INC/THE	COM	05969A105	30	1,448	SH		OTR	49	0	1,448	0
BANCORPSOUTH BANK	COM	05971J102	21,842	672,488	SH		DFND	15	583,171	0	89,317
BANCORPSOUTH BANK	COM	05971J102	150	4,617	SH		OTR	15,119	4,617	0	0
BANCORPSOUTH BANK	COM	05971J102	3,322	102,281	SH		DFND	22	102,281	0	0
BANCORPSOUTH BANK	COM	05971J102	3	90	SH		DFND	4	90	0	0
BANCORPSOUTH BANK	COM	05971J102	14,002	431,086	SH		DFND	48	431,086	0	0
BANCORPSOUTH BANK	COM	05971J102	1,780	54,805	SH		DFND	49	54,525	0	280
BANCORPSOUTH BANK	COM	05971J102	3	77	SH		OTR	49	0	77	0
BANCORPSOUTH BANK	COM	05971J102	0	3	SH		DFND	59	3	0	0
BANCORPSOUTH BANK	COM	05971J102	47	1,458	SH		DFND		1,458	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	24,342	192,065	SH		DFND	15	156,725	0	35,340
BANDWIDTH INC-CLASS A	COM	05988J103	126,270	996,291	SH		DFND	15,22	996,291	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	9,331	73,624	SH		DFND	22	73,624	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	462	3,645	SH		OTR	22,104	3,645	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	13	100	SH		DFND	4	100	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	2,811	22,179	SH		DFND	42	22,179	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	12,211	96,344	SH		DFND	48	96,344	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	940	7,419	SH		DFND	49	7,419	0	0
BANDWIDTH INC-CLASS A	COM	05988J103	195	1,541	SH		OTR	49	1,316	225	0
BANK FIRST CORP	COM	06211J100	1,107	14,758	SH		DFND	15	12,674	0	2,084
BANK FIRST CORP	COM	06211J100	1,170	15,605	SH		DFND	48	15,605	0	0
BANK OF AMERICA CORP	COM	060505104	26,448	683,598	SH		DFND	15,22	683,598	0	0
BANK OF AMERICA CORP	COM	060505104	26,963	696,895	SH		OTR	15,118	696,895	0	0
BANK OF AMERICA CORP	COM	060505104	14,747	381,150	SH		OTR	15,119	381,150	0	0
BANK OF AMERICA CORP	COM	060505104	1,276,350	32,989,143	SH		DFND	15	19,814,854	0	13,174,289
BANK OF AMERICA CORP	COM	060505104	139,971	3,617,765	SH		DFND	22	3,617,765	0	0
BANK OF AMERICA CORP	COM	060505104	7,498	193,807	SH		DFND	4	193,807	0	0
BANK OF AMERICA CORP	COM	060505104	4,105	106,094	SH		OTR	4	14,170	91,564	360
BANK OF AMERICA CORP	COM	060505104	325	8,390	SH		DFND	42	8,390	0	0
BANK OF AMERICA CORP	COM	060505104	891,786	23,049,525	SH		DFND	48	20,707,197	0	2,342,328
BANK OF AMERICA CORP	COM	060505104	36,853	952,509	SH		OTR	49	696,415	247,614	8,480
BANK OF AMERICA CORP	COM	060505104	322,402	8,332,953	SH		DFND	49	8,238,818	2,370	91,765
BANK OF AMERICA CORP	COM	060505104	11,333	292,924	SH		DFND	59	287,735	0	5,189
BANK OF COMMERCE HOLDINGS	COM	06424J103	424	33,234	SH		DFND	15	23,170	0	10,064
BANK OF COMMERCE HOLDINGS	COM	06424J103	477	37,423	SH		DFND	48	37,423	0	0
BANK OF COMMERCE HOLDINGS	COM	06424J103	38	3,000	SH		OTR	49	0	3,000	0
BANK OF HAWAII CORP	COM	062540109	154	1,719	SH		OTR	15,119	1,719	0	0
BANK OF HAWAII CORP	COM	062540109	23,050	257,576	SH		DFND	15	237,997	0	19,579
BANK OF HAWAII CORP	COM	062540109	3,825	42,737	SH		DFND	22	42,737	0	0
BANK OF HAWAII CORP	COM	062540109	23	260	SH		DFND	4	260	0	0
BANK OF HAWAII CORP	COM	062540109	13,770	153,870	SH		DFND	48	141,972	0	11,898
BANK OF HAWAII CORP	COM	062540109	2,244	25,074	SH		DFND	49	24,954	0	120
BANK OF HAWAII CORP	COM	062540109	7	77	SH		OTR	49	6	71	0
BANK OF HAWAII CORP	COM	062540109	676	7,556	SH		DFND	59	7,556	0	0
BANK OF HAWAII CORP	COM	062540109	0	1	SH		DFND		1	0	0
BANK OF MARIN BANCORP/CA	COM	063425102	1,646	42,037	SH		DFND	15	30,722	0	11,315
BANK OF MARIN BANCORP/CA	COM	063425102	1,320	33,701	SH		DFND	48	33,701	0	0
BANK OF MARIN BANCORP/CA	COM	063425102	14	360	SH		DFND	49	360	0	0
BANK OF MARIN BANCORP/CA	COM	063425102	0	1	SH		DFND		1	0	0
BANK OF MONTREAL	COM	063671101	492	5,518	SH		OTR	15,119	5,518	0	0
BANK OF MONTREAL	COM	063671101	39,809	446,689	SH		DFND	15	424,362	0	22,327
BANK OF MONTREAL	COM	063671101	29,406	329,959	SH		DFND	48	329,959	0	0
BANK OF MONTREAL	COM	063671101	305	3,417	SH		DFND	49	3,417	0	0
BANK OF MONTREAL	COM	063671101	404	4,530	SH		OTR	49	4,500	30	0
BANK OF MONTREAL	COM	063671101	154	1,733	SH		DFND	59	1,733	0	0
BANK OF MONTREAL	COM	063671101	0	1	SH		DFND		1	0	0
BANK OF N.T. BUTTERFIELD&SON	SHS	G0772R208	2,209	57,806	SH		DFND	15	13,163	0	44,643
BANK OF N.T. BUTTERFIELD&SON	SHS	G0772R208	4	116	SH		OTR	4	0	0	116
BANK OF N.T. BUTTERFIELD&SON	SHS	G0772R208	3,602	94,252	SH		DFND	48	94,252	0	0
BANK OF N.T. BUTTERFIELD&SON	SHS	G0772R208	197	5,150	SH		DFND	49	5,150	0	0
BANK OF N.T. BUTTERFIELD&SON	SHS	G0772R208	5	120	SH		DFND	59	120	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	2,564	54,222	SH		OTR	15,119	54,222	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	3,426	72,444	SH		DFND	15,22	72,444	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	1,234	26,093	SH		OTR	15,118	26,093	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	155,412	3,286,371	SH		DFND	15	1,951,949	0	1,334,422
BANK OF NEW YORK MELLON CORP	COM	064058100	6,121	129,430	SH		DFND	22	129,430	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	103,895	2,196,977	SH		DFND	48	1,955,390	0	241,587
BANK OF NEW YORK MELLON CORP	COM	064058100	1,959	41,417	SH		OTR	49	37,605	3,812	0
BANK OF NEW YORK MELLON CORP	COM	064058100	2,932	61,995	SH		DFND	49	61,995	0	0
BANK OF NEW YORK MELLON CORP	COM	064058100	75	1,592	SH		DFND	59	438	0	1,154
BANK OF NEW YORK MELLON CORP	COM	064058100	0	1	SH		DFND		1	0	0
BANK OF NOVA SCOTIA	COM	064149107	47,120	753,313	SH		DFND	15	711,254	0	42,059
BANK OF NOVA SCOTIA	COM	064149107	651	10,403	SH		OTR	15,119	10,403	0	0
BANK OF NOVA SCOTIA	COM	064149107	38,899	621,889	SH		DFND	48	621,889	0	0
BANK OF NOVA SCOTIA	COM	064149107	13	200	SH		DFND	49	200	0	0
BANK OF NOVA SCOTIA	COM	064149107	4	64	SH		DFND	59	64	0	0
BANK OZK	COM	06417N103	252	6,174	SH		OTR	15,119	6,174	0	0
BANK OZK	COM	06417N103	31,505	771,231	SH		DFND	15	716,138	0	55,093
BANK OZK	COM	06417N103	1,252	30,653	SH		OTR	22,87	30,653	0	0
BANK OZK	COM	06417N103	5,244	128,366	SH		DFND	22	128,366	0	0
BANK OZK	COM	06417N103	19	470	SH		DFND	4	470	0	0
BANK OZK	COM	06417N103	18,944	463,737	SH		DFND	48	428,442	0	35,295
BANK OZK	COM	06417N103	1,166	28,546	SH		DFND	49	28,316	0	230
BANK OZK	COM	06417N103	1	20	SH		OTR	49	20	0	0
BANK OZK	COM	06417N103	420	10,276	SH		DFND	59	10,276	0	0
BANK OZK	COM	06417N103	0	1	SH		DFND		1	0	0
BANK7 CORP	COM	06652N107	487	27,662	SH		DFND	15	26,477	0	1,185
BANK7 CORP	COM	06652N107	118	6,697	SH		DFND	48	6,697	0	0
BANKFINANCIAL CORP	COM	06643P104	1,092	105,766	SH		DFND	15	92,495	0	13,271
BANKFINANCIAL CORP	COM	06643P104	337	32,671	SH		DFND	48	32,671	0	0
BANKFINANCIAL CORP	COM	06643P104	24	2,300	SH		DFND	49	2,300	0	0
BANKUNITED INC	COM	06652K103	31,239	710,787	SH		DFND	15	630,584	0	80,203
BANKUNITED INC	COM	06652K103	214	4,873	SH		OTR	15,119	4,873	0	0
BANKUNITED INC	COM	06652K103	29,600	673,497	SH		DFND	22	673,497	0	0
BANKUNITED INC	COM	06652K103	1,125	25,599	SH		OTR	22,69	25,599	0	0
BANKUNITED INC	COM	06652K103	239	5,432	SH		OTR	22,87	5,432	0	0
BANKUNITED INC	COM	06652K103	85	1,945	SH		DFND	4	1,945	0	0
BANKUNITED INC	COM	06652K103	12	279	SH		OTR	4	0	0	279
BANKUNITED INC	COM	06652K103	12,403	282,209	SH		DFND	48	282,209	0	0
BANKUNITED INC	COM	06652K103	1,769	40,256	SH		DFND	49	40,256	0	0
BANKUNITED INC	COM	06652K103	49	1,117	SH		OTR	49	1,117	0	0
BANKUNITED INC	COM	06652K103	0	3	SH		DFND	59	3	0	0
BANKWELL FINANCIAL GROUP INC	COM	06654A103	644	23,892	SH		DFND	15	19,451	0	4,441
BANKWELL FINANCIAL GROUP INC	COM	06654A103	561	20,817	SH		DFND	48	20,817	0	0
BANKWELL FINANCIAL GROUP INC	COM	06654A103	0	1	SH		DFND		1	0	0
BANNER CORPORATION	COM	06652V208	21,852	409,743	SH		DFND	15	345,167	0	64,576
BANNER CORPORATION	COM	06652V208	5,889	110,432	SH		OTR	15,66	110,432	0	0
BANNER CORPORATION	COM	06652V208	3,741	70,141	SH		OTR	22,96	70,141	0	0
BANNER CORPORATION	COM	06652V208	8,436	158,193	SH		DFND	22	158,193	0	0
BANNER CORPORATION	COM	06652V208	108	2,025	SH		DFND	4	2,025	0	0
BANNER CORPORATION	COM	06652V208	5,078	95,224	SH		DFND	48	95,224	0	0
BANNER CORPORATION	COM	06652V208	2,538	47,585	SH		DFND	49	47,585	0	0
BAOZUN INC-SPN ADR	SPON	06684L103	1,243	32,602	SH		DFND	15	31,629	0	973
BAOZUN INC-SPN ADR	SPON	06684L103	2,007	52,618	SH		DFND	48	52,618	0	0
BAR HARBOR BANKSHARES	COM	066849100	1,631	55,441	SH		DFND	15	42,533	0	12,908
BAR HARBOR BANKSHARES	COM	066849100	1,241	42,198	SH		DFND	48	42,198	0	0
BARCLAYS PLC-SPONS ADR	ADR	06738E204	46	4,494	SH		DFND	4	4,494	0	0
BARCLAYS PLC-SPONS ADR	ADR	06738E204	14,329	1,400,658	SH		DFND	42	1,400,658	0	0
BARCLAYS PLC-SPONS ADR	ADR	06738E204	4,903	479,232	SH		DFND	49	479,232	0	0
BARCLAYS PLC-SPONS ADR	ADR	06738E204	186	18,215	SH		OTR	49	5,424	12,791	0
BARCLAYS PLC-SPONS ADR	ADR	06738E204	287	28,100	SH		DFND	59	28,094	0	6
BARCLAYS PLC-SPONS ADR	ADR	06738E204	6	545	SH		DFND		545	0	0
BARNES & NOBLE EDUCATION INC	COM	06777U101	789	96,941	SH		DFND	15	96,941	0	0
BARNES & NOBLE EDUCATION INC	COM	06777U101	562	69,073	SH		DFND	22	69,073	0	0
BARNES & NOBLE EDUCATION INC	COM	06777U101	130	15,957	SH		DFND	48	15,957	0	0
BARNES & NOBLE EDUCATION INC	COM	06777U101	0	1	SH		DFND		1	0	0
BARNES GROUP INC	COM	067806109	16,831	339,747	SH		DFND	15	297,637	0	42,110
BARNES GROUP INC	COM	067806109	5,969	120,489	SH		DFND	22	120,489	0	0
BARNES GROUP INC	COM	067806109	1,165	23,523	SH		OTR	22,71	23,523	0	0
BARNES GROUP INC	COM	067806109	24	480	SH		DFND	4	480	0	0
BARNES GROUP INC	COM	067806109	5,889	118,881	SH		DFND	48	118,881	0	0
BARNES GROUP INC	COM	067806109	2,079	41,973	SH		DFND	49	41,973	0	0
BARNES GROUP INC	COM	067806109	0	1	SH		DFND		1	0	0
BARRETT BUSINESS SVCS INC	COM	068463108	2,866	41,625	SH		DFND	15	35,766	0	5,859
BARRETT BUSINESS SVCS INC	COM	068463108	1,281	18,606	SH		DFND	48	18,606	0	0
BARRETT BUSINESS SVCS INC	COM	068463108	49	715	SH		DFND	49	715	0	0
BARRETT BUSINESS SVCS INC	COM	068463108	1	11	SH		DFND		11	0	0
BARRICK GOLD CORP	COM	067901108	304	15,333	SH		OTR	15,119	15,333	0	0
BARRICK GOLD CORP	COM	067901108	40,495	2,045,200	SH		DFND	15	1,916,925	0	128,275
BARRICK GOLD CORP	COM	067901108	9,696	489,703	SH		DFND	22	489,703	0	0
BARRICK GOLD CORP	COM	067901108	839	42,375	SH		OTR	40,48	42,375	0	0
BARRICK GOLD CORP	COM	067901108	5,776	291,715	SH		DFND	40,22	291,715	0	0
BARRICK GOLD CORP	COM	067901108	26,120	1,319,177	SH		DFND	40	1,319,177	0	0
BARRICK GOLD CORP	COM	067901108	18,067	912,484	SH		DFND	48	912,484	0	0
BARRICK GOLD CORP	COM	067901108	37	1,873	SH		OTR	49	107	1,766	0
BARRICK GOLD CORP	COM	067901108	51	2,587	SH		DFND	49	2,587	0	0
BARRICK GOLD CORP	COM	067901108	365	18,412	SH		DFND	59	17,389	0	1,023
BASSETT FURNITURE INDS	COM	070203104	808	33,308	SH		DFND	15	33,308	0	0
BASSETT FURNITURE INDS	COM	070203104	81	3,325	SH		DFND	48	3,325	0	0
BASSETT FURNITURE INDS	COM	070203104	1	58	SH		DFND	49	58	0	0
BAUDAX BIO INC	COM	07160F107	131	101,229	SH		DFND	48	101,229	0	0
BAUSCH HEALTH COS INC	COM	071734107	90	2,823	SH		OTR	15,119	2,823	0	0
BAUSCH HEALTH COS INC	COM	071734107	7,197	226,763	SH		DFND	15	215,395	0	11,368
BAUSCH HEALTH COS INC	COM	071734107	5,197	163,739	SH		DFND	48	163,739	0	0
BAUSCH HEALTH COS INC	COM	071734107	1,704	53,685	SH		OTR	49	237	53,448	0
BAUSCH HEALTH COS INC	COM	071734107	2	62	SH		DFND	49	62	0	0
BAUSCH HEALTH COS INC	COM	071734107	16	505	SH		DFND	59	505	0	0
BAUSCH HEALTH COS INC	COM	071734107	0	1	SH		DFND		1	0	0
BAXTER INTERNATIONAL INC	COM	071813109	1,437	17,036	SH		OTR	15,118	17,036	0	0
BAXTER INTERNATIONAL INC	COM	071813109	191,954	2,275,949	SH		DFND	15	1,419,605	0	856,344
BAXTER INTERNATIONAL INC	COM	071813109	1,025	12,154	SH		OTR	15,119	12,154	0	0
BAXTER INTERNATIONAL INC	COM	071813109	3,825	45,353	SH		DFND	15,22	45,353	0	0
BAXTER INTERNATIONAL INC	COM	071813109	7,043	83,513	SH		DFND	22	83,513	0	0
BAXTER INTERNATIONAL INC	COM	071813109	820	9,726	SH		DFND	4	9,726	0	0
BAXTER INTERNATIONAL INC	COM	071813109	141,599	1,678,910	SH		DFND	48	1,523,509	0	155,401
BAXTER INTERNATIONAL INC	COM	071813109	4,367	51,776	SH		OTR	49	9,443	41,835	498
BAXTER INTERNATIONAL INC	COM	071813109	20,320	240,924	SH		DFND	49	240,308	0	616
BAXTER INTERNATIONAL INC	COM	071813109	108	1,279	SH		DFND	59	1,092	0	187
BAXTER INTERNATIONAL INC	COM	071813109	51	609	SH		DFND		609	0	0
BAYCOM CORP	COM	07272M107	371	20,601	SH		DFND	15	14,825	0	5,776
BAYCOM CORP	COM	07272M107	449	24,902	SH		DFND	48	24,902	0	0
BCB BANCORP INC	COM	055298103	846	61,283	SH		DFND	15	52,313	0	8,970
BCB BANCORP INC	COM	055298103	539	39,057	SH		DFND	48	39,057	0	0
BCE INC	COM	05534B760	60	1,326	SH		OTR	15,119	1,326	0	0
BCE INC	COM	05534B760	16,048	355,506	SH		DFND	15	350,366	0	5,140
BCE INC	COM	05534B760	3,560	78,861	SH		DFND	48	78,861	0	0
BCE INC	COM	05534B760	311	6,897	SH		OTR	49	164	6,733	0
BCE INC	COM	05534B760	272	6,020	SH		DFND	49	6,020	0	0
BCE INC	COM	05534B760	4,731	104,812	SH		DFND	59	102,318	0	2,494
BCE INC	COM	05534B760	0	1	SH		DFND		1	0	0
BEACON ROOFING SUPPLY INC	COM	073685109	213	4,077	SH		OTR	15,119	4,077	0	0
BEACON ROOFING SUPPLY INC	COM	073685109	5,386	102,952	SH		DFND	15	55,931	0	47,021
BEACON ROOFING SUPPLY INC	COM	073685109	1,688	32,259	SH		OTR	22,86	32,259	0	0
BEACON ROOFING SUPPLY INC	COM	073685109	7,269	138,928	SH		DFND	48	138,928	0	0
BEACON ROOFING SUPPLY INC	COM	073685109	581	11,099	SH		OTR	49	0	11,099	0
BEACON ROOFING SUPPLY INC	COM	073685109	4	75	SH		DFND	49	75	0	0
BEACON ROOFING SUPPLY INC	COM	073685109	15	288	SH		DFND	59	288	0	0
BEAM THERAPEUTICS INC	COM	07373V105	4,205	52,535	SH		DFND	15	38,711	0	13,824
BEAM THERAPEUTICS INC	COM	07373V105	1,914

Bank Of New York Mellon (BK) 13F-HRQuarterly holdings report by institutional manager